Difference?
A fish owner asked the question, "I was told that for
my community tank I should get fish that are
omnivores. What are omnivore, herbivore and
carnivore, and what is the difference? "
Answer: You've posed an excellent question that all
too few people concern themselves with.Not all fish
require the same diet. Like other organisms, a fish is
designed with a mouth, teeth, and an digestive tract
that is intended for certain types of food. It's
important to keep in mind that any living creature will
eat virtually anything if they are hungry enough. In
other words, don't make broad assumptions about the
dietary needs of a fish based on observation alone.Do
your homework and find out what the fish needs to
eat to remain healthy. There are three basic categories
that fish can be classified in, based on their dietary
needs. Those are: carnivore, herbivore, and omnivore.
Carnivores are meat eaters, generally requiring live
foods. They have a large mouth with sharp pointed
teeth that allow them to grasp their prey and tear off
large chunks of flesh, which is swallowed whole rather
than ground or chewed first.Carnivores have a short
intestinal tract, and a relatively large stomach
designed to hold an entire fish. Their digestive system
lacks the ability to digest vegetable matter, so even
though they might eat plants, they cannot derive
nutrients from them as other fish do. Because they will
chase down and eat other fish in the aquarium,
carnivores are not suitable for a community tank.
Herbivores are on the opposite end of the dietary
Life
The coral reef environment has a physical
structure that is radically different than that of
the surrounding open ocean. Fishes that live in
reef habitats have therefore evolved a host of
specialized traits well suited to their colorful
and complex homes.
Warning coloration
Parrotfish beak
Herbivores
Herbivorous fishes are those that feed
mainly or entirely on plant material.
Planktivores
Planktivorous coral reef fishes are those that
prey upon small animal plankton
(zooplankton).
Open Water Plankton Feeders: The daytime openwater plankton feeding "fraternity" of coral reef fish
consists of a hodgepodge of species of diverse
heritage, including damselfishes, wrasses, snappers,
sea basses, and surgeonfishes.
Benthic Carnivores
Piscivores
Piscivorous coral reef fish are those that prey
mainly or entirely upon other fishes.
There are three different basic hunting
strategies employed by such predators (see
below).
Introduction
Fishes living in the Caribbean Sea rely on a variety of food
sources including plants, plankton, invertebrates, and other
fishes. Fishes can feed either on the reef or off of the reef in
the sandy bottoms or sea grass beds. Fishes can feed either
diurnally (during the day) or nocturnally (at night). Their diet
and mode of feeding strongly influences their morphology.
Moreover, their foraging strategy should affect their
susceptibility to predators and thus influence their antipredator mechanisms.
Herbivorous Fishes
Only 10 25% of species on a Caribbean reef fishes are
herbivores. Although there are relatively few herbivorous fish
species, these species tend to have relatively large population
sizes such that the total biomass of herbivores is high.
Herbivorous fishes are most common in shallow water which
is not surprising because light levels are higher in shallow
water which promotes algal growth. In general, herbivores are
the most common fishes above 30 feet whereas carnivores are
the most common fishes below 30 feet. On shallow reefs
herbivores may take over 150,000 feeding bites per meter
each day and consume from half to all of the daily algal
production (one reason that there is limited algal growth on a
healthy reefs).
Nomadic Herbivores
Sedentary Feeders
Planktivorous Fishes
Most reef fishes eat plankton during their juvenile stage and
some continue to feed on plankton after they have matured.
Planktivores feed either above the reef or feed near the reef or
sandy bottom and may feed during the day or at night.
Diurnal planktivorous fishes that feed above the reef are able
to capture the plankton before it reaches the species living on
the reef. Some of these species are very large (e.g., Giant
Manta (Manta birostris) and Whale Sharks (Rhincodon
typus)) and swim through the water column with their mouths
open picking up plankton. Because they are so large, they
generally do not need to worry about predators. These species
often forage alone.
Carnivorous Fishes
Carnivorous fishes make up from 41 - 68% of species on the
reef.
Species that feed off of the reef face some problems including
greater risk of predation, finding prey that are hidden beneath
the sand, and dealing with prey protected by large shells.
Thus, relatively few species feed on invertebrates off of the
reef. Species feeding in this way use a variety of strategies.
Spotted Eagle Rays (Aetobatus narinari) feed on clams,
oysters, and gastropods that are buried beneath the sand.
They crush the shells of their prey using specialized dental
plates. Goatfishes (Family Mullidae) feed in the sand, but
generally remain near the safety of the reef. They have
appendages near their chins, barbells, which they use to detect
prey by touch and taste. Trunkfishes are so well protected
from predators that they are able to move far from the safety
of the reef. When they identify signs of prey, they uncover the
prey by shooting a jet of water out of their mouths. Hogfishes
(Family Labridae) are large enough that they are safe from
most predators. They have large jaws and teeth that allow
them to feed on well-protected prey.
At night, invertebrates come out to feed on the sand flats and
in sea grass beds. Thus, there is a lot of food for invertebratefeeders at night off of the reef. Because it is dark, predators
can not rely on vision to detect their prey. Instead they rely on
touch, taste, smell, and their lateral lines. Because these
senses are not as effective as sight in determining the exact
location of a prey item, they have had to come up with
Moray eels (Family Muranidae) may hunt for fishes and other
prey at night either in the open or by using their long narrow
bodies to travel into the protected crevices in the reef where
other fishes are sleeping.
Stalking predators begin their attacks from a stationary
position. They attempt to maneuver themselves into position
and wait for prey to approach close enough to attack. Their
bodies are thin and elongated in order to reduce the area
exposed to prey. Their tails have large surface area needed to
provide the power necessary for a rapid strike. Trumpetfish
(Aulostomus maculates) are classic stalking predators found
on coral reefs. They use their expandable jaws to suck in
fishes. They are capable of swallowing unexpectedly large prey
so it is possible to observe a bulge in the stomach of a fish who
has recently fed. Trumpetfish often orient themselves
vertically among sea fans or soft corals. Foraging trumpetfish
may make 2-3 strikes per hour. Researchers have observed 8
successful strikes out of 45 attempts in 19 hours of
investigation.
Ambush predators wait for prey to get close to them and then
they grab them. Ambush predators are often well camouflaged
and possess upturned mouths that they use to capture their
prey. For example, the foraging strategy of groupers (Family
Serranidae) is to sit, wait, and inhale. They capture their prey
by the suction created by the rapid expansion of their jaws and
the flaring of their gill covers. Nassau Groupers (Epinephelus
striatus) tend to be somewhat territorial and can live in the
same home range for a number of years. Their diet includes
fishes, including parrotfish, grunts, and snappers, lobsters,
and mollusks.
Fish make up 80% of the diet of Grasbys (Cephalopholis
cruentatus). Their favorite food is Brown Chromis, but they
also feed on Bicolor Damselfish (Stegastes partitus), Blue
Chromis, and Yellow Goatfish (Mulloidchthys martinicus).
The remainder of their diet consists of invertebrates, mostly
crustaceans. Coneys (Cephalopholis fulvus) prefer to live on
isolated patch reefs. Their diet is made up of an equal mix of
small fishes and invertebrates.
Flounders (Family Bothidae) are the master of camouflage.
They can change colors to match their surroundings and lie in
wait until and unsuspecting prey passes by. Other ambush
predators include scorpionfishes (Family Scorpaenidae),
toadfishes (Family Batrachoididae), and lizardfishes (Family
Synodontidae).
MarineEcosystems
Like all ecosystems, marine ecosystems are mostly self-sustaining systems of life forms and the physical environment. In these ecosystems mate
have certain things in common and marine ecosystems have a few unique twists to these cycles. We can begin this discussion with describing the mak
Baby two spot octopus showing chromatophore cells (in orange and black) that it can use to change color. These cells cover the skin of m
and come in a variety of colors - both orange and black are seen here. The pigment in these cells can be stretched to create different ski
octopus can either stand out against the background -or- blend in remarkably well. Image by Shane Anderson
Almost all life forms are made of a cell (if they are unicellular) or cells (if they are multicellular) and the products of the cell(s). To sustain life, these
chemical reaction called cellular respiration.
Cellular respiration is the life sustaining process for all life forms, including both plant and animal types. It is where a cell uses what we call 'cell fo
carbohydrate molecule - usually glucose made of six carbons, twelve hydrogens, and six oxygens) in the presence of oxygen to make the 'cell energy' (
molecule we call ATP) and with a waste product of carbon dioxide. The ATP is used to repair the cell and keeps it alive. A few organisms can cellularly r
but it is not as efficient and we call these critters 'anaerobes.' Most critters need oxygen to stay alive and 'cell food.' The cell food is obtained by eating i
consumers) and by photosynthesis in critters we call plants (also called producers).
Photosynthesis equation.
Photosynthesis is the food-making process for producers, which we often call plants. In the marine environment many of the producers are micros
kingdom called Protoctista, along with the algae. But, they are still like plants in that they photosynthesize no matter what group they are classified as. P
carbon dioxide and water are combined within a specialized part of a cell (called a chloroplast), in the presence of light, to form the 'cell food' molecule c
a waste product of oxygen. In a sense it is cellular respiration in reverse but it will only happen in cells with chloroplasts (in which there is the molecule c
involved). It is a complicated reaction but the end product provides the recycling of carbon dioxide back to oxygen for the cycle of these two gases. With
life forms (plants and animals) would use up the oxygen on Earth and it would only be found as carbon dioxide (not usable in cellular respiration).
Multicellular adult two spot octopus individual with most of its orange chromatophores turned on so that its skin appears orange. Notice th
where blue chromatophore cells are turned on making spots that appear as large eyes and thus its common name. Its real eyes are just abo
Image by Genny Anderson
Both unicellular (one celled) and multicellular (more than one cell) individuals are found in the marine ecosystem taking on various levels of inter
unicellular plants and animals. There are multicellular plants and animals as well. The multicellular species may be very simple or more complex with ce
tissues, organs, and organ systems depending on the species. Whether or not a species is simple or complex they each have their place in the ecosyst
stories about their life histories.
Two (or more) two spot octopods in an area would be a population. Notice that these individuals have turned off most of the orange chromat
pattern on their skin that almost matches their environment - octopus are very good at blending in due to their chromatophore skin cells
Anderson
Populations are groups of organisms of one species living in an area and interacting. Interactions between individuals of the same species are term
interactions and can be positive (like cooperation), negative (like competition) or anything in between.
A simple community associated with the two spot octopus (seen above, left, out hunting in the tidepools at low tide) would include its food
the kelp crab (kelp, right). In reality the community would be more complex including a variety of food items for the octopus (fish, clams) a
Images by Genny Anderson
Communities are composed of populations of many species living in an area and interacting. Interactions between species are termed interspecific
interactions, can be positive, negative or anything in between.
Tidepool habitats of the two spot octopus include many species interacting in a complex community. It could even be considered a small eco
of the interactions occur here and it is pretty much self-sustaining (although the tide brings in new species and nutrients with each cycle
Anderson.
Ecosystems are areas where the community, or communities, are rather self-sustaining. This involves what we call food chains and webs - that i
who and following the energy flow (in the form of food) all the way to what we call the bottom of the food chain. The bottom is where the producers are f
are the photosynthesizers and depend on carbon dioxide and light so that they can make their own food. Gases are cycled and recycled in ecosystems
respiration and photosynthesis. Nutrients are cycled by decomposers (bacteria and fungi) in ecosystems that are involved with rotting (decomposing) an
releasing the nutrients from these cells. Food is cycled through what we call trophic pyramids - always with the producers at the base. Ecosystems can
large areas (like the entire planet Earth) or smaller subunits (like a tidepool habitat shown here) depending on how strict one uses the term 'self-sustain
The trophic pyramid of an ecosystem. Kelp could be placed on the bottom as a producer, the kelp crab on the middle layer (as a primary co
and the two spot octopus on the top (as a secondary consumer or carnivore).
Trophic pyramids in the ocean are similar to those on land with the base always larger than the upper trophic levels. The base is the producers. Th
level of the critters that eat the producers (we call these herbivores, or plant eating animals). The third level is composed of critters that eat the herbivor
carnivores, or animal eating animals). As you move between the levels there is always a loss of biomass because the transfer of energy is usually only
This means that if there were 100 pounds of producers in an area there would be enough food for 10 pounds of herbivores but the 10 pounds of herbivo
1 pound of a carnivore. So you see, as you move up the food chain (or trophic pyramid) there is a smaller and smaller biomass that can be supported. S
the second and third levels of the trophic pyramid as primary consumers (the herbivores) and secondary consumers (the first order carnivores). There c
(each with a loss of 90 percent biomass average) with tertiary consumers (carnivores eating the secondary consumers) and so on.
Marine producers are either seaweeds and/or phytoplankton and must be found where there is enough light for photosynthesis. Therefore the mari
the ocean, the area we call the photic zone. Animals can be found everywhere, in both the photic and aphotic zones, because they are not limited by lig
ocean bottom so their distribution is limited to the edges of continents and islands where the depth is within the photic zone. These seaweeds account f
ocean because they are so geographically limited. Most of the ocean is open ocean, away from the edges of land and over deep water. It is here, near
plankton) dominates. Phytoplankton is also found in the same coastal areas as the seaweeds. Both types of producers are important in marine ecosyst
ecosystems are more common. The importance of the phytoplankton is one of the things that make marine ecosystems unique and different from land e
generally microscopic. Land based ecosystems are based primarily on large land plants but marine ecosystems are based primarily on tiny, microscopi