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BIOENERGTICA
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BIOENERGTICA
Definicin:
Es el anlisis cuantitativo de las transformaciones
y utilizacin de la energa en los sistemas biolgicos.
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BIOENERGTICA
Definicin:
Es el anlisis cuantitativo de las transformaciones
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BIOENERGTICA
Definicin:
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BIOENERGTICA
Definicin:
Es el anlisis cuantitativo de las transformaciones
y utilizacin de la energa en los sistemas biolgicos.
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Clasificacin de los organismos vivos segn
la forma de obtener energa
Fottrofos
Obtienen energa
de la luz solar
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Clasificacin de los organismos vivos segn
la forma de obtener energa
C6H12O6
Ej. Glucosa
Comp. orgnicos
Fottrofos
Obtienen energa
de la luz solar
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Clasificacin de los organismos vivos segn
la forma de obtener energa
C6H12O6
Ej. Glucosa
Comp. orgnicos
Fottrofos Quimitrofos
Obtienen energa Utilizan la energa qumica obtenida
de la luz solar de compuestos orgnicos
(nutrientes).
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BIOENERGTICA
Definicin:
Es el anlisis cuantitativo de las transformaciones
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BIOENERGTICA
Definicin:
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BIOENERGTICA
Definicin:
Es el anlisis cuantitativo de las transformaciones
y utilizacin de la energa en los sistemas biolgicos.
obedecen
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Cules son las condiciones bajo las
que se producen en forma espontnea
los procesos vitales?
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Las Leyes de la termodinmica
Primera ley
(principio de conservacin de la energa)
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Las Leyes de la termodinmica
Primera ley
(principio de conservacin de la energa)
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Las Leyes de la termodinmica
Primera ley
(principio de conservacin de la energa)
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Universo = Sistema + Entorno
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Universo = Sistema + Entorno
Sistema: parte del universo que elegimos
estudiar; es el conjunto de materia que
est experimentando un proceso qumico
o fsico determinado.
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Universo = Sistema + Entorno
Sistema: parte del universo que elegimos
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est experimentando un proceso qumico
o fsico determinado.
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Universo = Sistema + Entorno
Sistemas:
Abierto:
Cerrado:
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Universo = Sistema + Entorno
Sistemas:
Abierto: intercambia Materia y Energa
Cerrado:
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Universo = Sistema + Entorno
Sistemas:
Abierto: intercambia Materia y Energa
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Universo = Sistema + Entorno
Sistemas:
Abierto: intercambia Materia y Energa
Nutrientes Animal desechos (desorden)
generan trabajo
y calor
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Universo = Sistema + Entorno
Sistemas:
Abierto: intercambia Materia y Energa
Nutrientes Animal desechos (desorden)
SERES VIVOS
NUNCA ESTAN EN generan trabajo
EQUILIBRIO
y calor
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SERES VIVOS
NUNCA ESTAN EN EQUILIBRIO
TERMODINMICO
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Los sistemas vivos se mantienen alejados del equilibrio termodinmico
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Estado estacionario dinmico: UN EJEMPLO
Alimentos GLU
(carbohidratos)
CO2
(deshecho)
CO2
(deshecho)
Depsito Grasa
Hgado
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Las Leyes de la termodinmica
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Las Leyes de la termodinmica
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Las Leyes de la termodinmica
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Las Leyes de la termodinmica
Tipos de trabajos:
Crecimiento
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La energa Trabajo
de Se transforma en Calor
nutrientes Desorden
Tipos de trabajos:
Espermatozoides
Crecimiento
Reproduccin
Produccin: Leche, Lana, Carne, Huevos
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La energa Trabajo
de Se transforma en Calor
nutrientes Desorden
Tipos de trabajos:
Espermatozoides
Crecimiento
Movimiento
Reproduccin
Produccin: Leche, Lana, Carne, Huevos
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ORDEN
La energa Trabajo
de nutrientes Se transforma en Calor
Desorden
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La energa Trabajo
de nutrientes Se transforma en Calor
Desorden
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La energa Trabajo
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Desorden
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La energa Trabajo
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Desorden
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La energa Trabajo
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Desorden
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
= G
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema.
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema.
A B S = +
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema.
A B S = +
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema. Proceso espontneo
A B S = +
+ complejos + simples
+ ordenados + desordenados Ganancia de entropa
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema. Proceso espontneo
A B S = +
+ complejos + simples
+ ordenados + desordenados Ganancia de entropa
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema.
A B S = +
+ complejos + simples
+ ordenados + desordenados Ganancia de entropa
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Entropa (S)
Expresin cuantitativa del desorden de un
sistema.
A B S = +
+ complejos + simples
+ ordenados + desordenados Ganancia de entropa
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Entalpa (H)
Es el contenido calrico del sistema
reaccionante:
Si libera calor:
Cuando la energa en forma de calor es cedida por el sistema
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Entalpa (H)
Es el contenido calrico del sistema
reaccionante:
Si libera calor: exotrmica (H < 0)
Cuando la energa en forma de calor es cedida por el sistema
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Entalpa (H)
Es el contenido calrico del sistema
reaccionante:
Si libera calor: exotrmica (H < 0)
Cuando la energa en forma de calor es cedida por el sistema
Si adquiere calor:
Es el calor ganado por el sistema
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Entalpa (H)
Es el contenido calrico del sistema
reaccionante:
Si libera calor: exotrmica (H < 0)
Cuando la energa en forma de calor es cedida por el sistema
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Entalpa (H)
Es el contenido calrico del sistema
reaccionante:
Si libera calor: exotrmica (H < 0)
Cuando la energa en forma de calor es cedida por el sistema
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
Es la energa que usan las clulas
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
Es la energa que usan las clulas
Predice:
1. La direccin de la reaccin qumica
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
Es la energa que usan las clulas
Predice:
1. La direccin de la reaccin qumica
2. Su posicin exacta de equilibrio
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
Es la energa que usan las clulas
Predice:
1. La direccin de la reaccin qumica
2. Su posicin exacta de equilibrio
3. La cantidad de trabajo desarrollada
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Energa libre de Gibbs (G)
Cantidad de energa capaz de realizar trabajo
durante una reaccin a presin y temperatura
cte.
Es la energa que usan las clulas
Predice:
1. La direccin de la reaccin qumica
2. Su posicin exacta de equilibrio
3. La cantidad de trabajo desarrollada
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los procesos vitales?
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Energa libre de Gibbs (G) 1878
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Energa libre de Gibbs (G) 1878
A B
G = GB - GA
- = reaccin exergnica (espontnea)
Pueden realizar trabajo
+ = reaccin endergnica
0 = proceso en equilibrio
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Estado inicial y final del sistema:
Variaciones ( )
C6H12O6 + 6 O2 6 CO2 + 6 H2O
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En sistemas biolgicos (T y P cte.), las variaciones de G, H y S
estn relacionadas por la ecuacin:
G = H - T S
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En sistemas biolgicos (T y P cte.), las variaciones de G, H y S
estn relacionadas por la ecuacin:
G = H - T S
Indica:
El gradiente de concentracin mxima que puede generar
La actividad muscular mxima que puede generar
La separacin mxima de carga que puede establecer, etc.
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G = H - T S 1 cal = 4.184 J
kJ/mol kJ/mol K kJ/mol.K
1. G
( - ) reaccin libera energia libre = exergnica
(+) reaccin gana energia libre = endergonica
2. H
( - ) reaccin libera calor = exotrmica
(+) reaccin gana calor = endotrmica
3. S
( + ) reaccin aumenta entropa (P > desordenados S)
( - ) reaccin disminuye entropa
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C6H12O6 + 6 O2 6 CO2 + 6 H2O
G = H - T S
- - +
Reaccin Reaccin Reaccin
exergnica exotrmica aumenta entropa
Reaccin es favorable,
sistema reacciona de forma espontnea
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6 CO2 + 6 H2O C6H12O6 + 6 O2
G = H - T S
+ + -
Reaccin Reaccin Reaccin
endergnica endotrmica disminuye entropa
Reaccin NO es favorable
Reaccin tiende a transcurrir en sentido inverso
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G = H - TS
Signos: - - +
Proceso favorable
2da Ley:
todo proceso que ocurre espontneamente es
capaz de realizar trabajo
G es una medida de la capacidad del sistema para producir
trabajo, sirve para determinar si una reaccin puede o no
ocurrir espontneamente
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La G depende de las condiciones
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La G depende de las condiciones
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La G depende de las condiciones
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G
G = G final - G inicial
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G
G=-RTlnKeq
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G
G = - R T ln Keq
En condiciones estndar:
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
pH biolgico = 7 [ H+] 10-7 M
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
pH biolgico = 7 [ H+] 10-7 M
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
pH biolgico = 7 [ H+] 10-7 M
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
pH biolgico = 7 [ H+] 10-7 M
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G
G = - R T ln Keq
En condiciones estndar:
reactivos y productos 1 M
1 atm
(temperatura absoluta) T = 298 K, en grados K
(25 C)
pH corregido: [H+] 1M pH=0
pH biolgico = 7 [ H+] 10-7 M
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial aA + bB cC + dD
Final
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial aA + bB cC + dD
Final
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial aA + bB cC + dD
Final
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial aA + bB cC + dD
Final
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial Glucosa 1-P Glucosa 6-P
1 1
G
Glucosa 1-P Glucosa 6-P
1 19
Final
Keq = G6P = 19 = 19
G1P 1
G = - (8.315 J/mol.K) (298 K) (ln 19)
G = - 7300 J/mol o -7.3 kJ/mol
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G = - R T ln Keq
En condiciones estndar:
Los reactivos y productos 1 M
Inicial Glucosa 1-P Glucosa 6-P
1 1
G
Glucosa 1-P Glucosa 6-P
1 19
Final G = - (8.315 J/mol.K) (298 K) (ln 19)
G = - 7300 J/mol o -7.3 kJ/mol
G = - RT ln Keq
aA + bB cC + dD
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
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a, b, c y d son el nmero de molculas de A, B, C y D que participan.
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G est directamente relacionada con la Keq
forma alternativa de
G = - RT ln Keq
expresar la Keq.
aA + bB cC + dD
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
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G est directamente relacionada con la Keq
forma alternativa de
G = - RT ln Keq
expresar la Keq.
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
Keq G Reaccin
K<1 Positiva (+) se favorece la formacin de reactivos.
K>1 Negativa (-) se favorece la formacin de productos
K=1 Cero se encuentra en equilibrio
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G est directamente relacionada con la Keq
forma alternativa de
G = - RT ln Keq
expresar la Keq.
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
Keq G Reaccin
K<1 Positiva (+) se favorece la formacin de reactivos.
K>1 Negativa (-) se favorece la formacin de productos
K=1 Cero se encuentra en equilibrio
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G est directamente relacionada con la Keq
forma alternativa de
G = - RT ln Keq
expresar la Keq.
Keq = [ C ]c [ D ]d
[ A ]a [ B ]b
Keq G Reaccin
K<1 Positiva (+) se favorece la formacin de reactivos.
K>1 Negativa (-) se favorece la formacin de productos
K=1 Cero se encuentra en equilibrio
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G
G = G final - G inicial
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G
VARIACION DE ENERGIA LIBRE REAL
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VARIACION DE ENERGIA LIBRE REAL, G
G = G + RT ln [C][D]
[A][B]
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VARIACION DE ENERGIA LIBRE REAL, G
G = G + RT ln [C][D]
[A][B]
Temperatura (K) en la cual
se desarrolla la reaccin
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VARIACION DE ENERGIA LIBRE REAL, G
G = G + RT ln [C][D] Q
[A][B]
Temperatura (K) en la cual Concentraciones Iniciales de Reacitvos
se desarrolla la reaccin y Productos en condiciones reales!
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VARIACION DE ENERGIA LIBRE REAL, G
G = G + RT ln [C][D] Q
[A][B]
Temperatura (K) en la cual Concentraciones Iniciales de Reacitvos
se desarrolla la reaccin y Productos en condiciones reales!
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VARIACION DE ENERGIA LIBRE REAL, G
G = G + RT ln [C][D] Q
[A][B]
Temperatura (K) en la cual Concentraciones Iniciales de Reacitvos
se desarrolla la reaccin y Productos en condiciones reales!
Temperatura
Depende:
Q
ESPONTANEIDAD DE UNA REACCIN = G
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Estado inicial
G = G + RT ln [C][D] Q
[A][B]
G = - RT ln Keq
G = 0 G = - RT ln Q
Keq
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Estado inicial
G = G + RT ln [C][D] Q
[A][B]
G = - RT ln Keq
G = 0 G = - RT ln Q
-G/RT
Keq= [C][D] = e Keq
[A][B]
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Espontaneidad de las reacciones
G = G + RT ln [C][D]
[A][B]
+
Puede una reaccin ocurrir espontneamente siendo su G +?
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Espontaneidad de las reacciones
G = G + RT ln [C][D]
[A][B]
+
Valor negativo y > G
-
Espontnea
G depende de:
la concentraciones de reactivos y
productos,
y de la temperatura durante la reaccin.
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G
Cantidad de energa capaz de realizar
trabajo
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G
Cantidad de energa capaz de realizar
trabajo
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G
Cantidad de energa capaz de realizar
trabajo
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G
Cantidad de energa capaz de realizar
trabajo
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Una reaccin termodinmicamente desfavorable
(endergnica) cmo puede ser impulsada en la
direccin directa?
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Una reaccin termodinmicamente desfavorable
(endergnica) cmo puede ser impulsada en la
direccin directa?
S, gracias al acoplamiento de reacciones
secuenciales a travs de un intermediario comn.
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Una reaccin termodinmicamente desfavorable
(endergnica) cmo puede ser impulsada en la
direccin directa?
S, gracias al acoplamiento de reacciones
secuenciales a travs de un intermediario comn.
1) A B G1
2) B C G2
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Una reaccin termodinmicamente desfavorable
(endergnica) cmo puede ser impulsada en la
direccin directa?
S, gracias al acoplamiento de reacciones
secuenciales a travs de un intermediario comn.
1) A B G1
Se suman
2) B C G2
A C G1 + G2
Gtotal
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Una reaccin termodinmicamente desfavorable
(endergnica) cmo puede ser impulsada en la
direccin directa?
S, gracias al acoplamiento de reacciones
secuenciales a travs de un intermediario comn.
1) A B G1
Se suman
2) B C G2
A C G1 + G2
Gtotal
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Glucosa+ Pi Glucosa 6-P + H2O G= 13.8 kJ/mol
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Glucosa+ Pi Glucosa 6-P + H2O G= 13.8 kJ/mol
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Glucosa+ Pi Glucosa 6-P + H2O G= 13.8 kJ/mol
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Las G son aditivas
Glucosa+ Pi Glucosa 6-P + H2O G= 13.8 kJ/mol
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Cual de las siguientes reacciones
es la ms rpida?
Reaccin G (Kj/mol)
1 - 13.4
2 -20.0
3 3.7
4 5.2
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Cual de las siguientes reacciones
es la ms rpida?
Reaccin G (Kj/mol)
1 - 13.4
2 -20.0
3 3.7
4 5.2
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Cual de las siguientes reacciones
es la ms rpida?
Reaccin
G (Kj/mol)
1 tema de la-cintica
La velocidad es un 13.4 qumica,
NO de la Termodinmica -20.0
2
3 3.7
4 5.2
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
G = Energa de activacin
Energa libre, G
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
GSP
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
GSP
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
GSP GPS
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
GSP GPS
Reactivo
Gi G
Producto
Gf
Coordenada de la reaccin
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
Estado de transicin G = Energa de activacin
Energa libre, G
GSP GPS
Inicial
Reactivo
G Gi G
Final Producto
Gf
Coordenada de la reaccin
la fuerza propulsora del
sistema hacia el equilibrio es:
G, es la fuerza motriz.
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Ej: combustin lea muy favorable termodinamicamente,
pero es estable ENERGIA DE ACTIVACION (G) es
superior a la disponible a temperatura ambiente.
Y en la CELULA?
COMO VENCEMOS ESTA G ???
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Ejercicios
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1) Cual es el G de hidrlisis del ATP?
ATP + H2O ADP + Pi
G = - RT ln Keq
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1) Cual es el G de hidrlisis del ATP?
ATP + H2O ADP + Pi
G = - RT ln Keq
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1) Cual es el G de hidrlisis del ATP?
G = - RT ln Keq
Glucosa 6-P + H2O Glucosa + Pi Keq= 270
G = - 8.135 x 10-3 kJ X 298 K X ln 270 = - 13.57 kJ/mol
mol.K
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1) Cual es el G de hidrlisis del ATP?
ATP + H2O ADP + Pi
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2) G de hidrlisis del ATP en glbulos rojos
G = G + RT ln [ADP][Pi]
[ATP]
G = - 30.500 J/mol+ (8.315 J/mol.K)(298 K)ln 1.83 X10-4
G = - 51.800 J/mol
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Resumen 1
Los organismos vivos crean y mantienen sus estructuras
complejas y ordenadas a expensas de la energa que
extraen de los combustibles.
En cualquier proceso fsico o qumico la cantidad de
energa total del universo permanece constante, pero
puede transformarse.
Las clulas son mquinas qumicas que funcionan a
temperatura y presin constantes, e intercambian energa
y materia con el entorno (nunca estan en equilibrio).
En todos los procesos la entropa del universo aumenta, el
orden de los seres vivos se obtiene a expensas del
desorden del entorno.
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Resumen 2
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Muchas gracias!!!
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No confundir
Equilibrios de reaccin asociados
a G (energa libre)
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Las Keq son multiplicativas
Glucosa+ Pi Glucosa 6-P + H2O Keq1= 3.9 x 10-3 M-1
G = - R T ln Keq
G = - 16.5 Kj/mol
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Clasificacin de los organismos vivos segn
la forma de obtener carbono
CO2
Comp. inorgnicos
H2O
O2
Ej. Glucosa
Comp. orgnicos
Auttrofos Hetetrofos
Autoalimentados Alimentados a partir de otros
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Reactivo Producto
Energa inicial (Gi) Energa final (Gf) G
G = Energa de activacin
Energa libre, G
Reactivo
Gi
Producto
Gf
Coordenada de la reaccin
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