Costello, Emblow and White (eds), 2001
HEMICHORDATA
Compiled by Jacob van der Land.
The European l ist was derived from URMO with
names provided by J. Hallan. No expert has checked
the accuracy of the ERMS list.
PHYLUM HEMICHORDATA
Class Enteropneusta
Family Harrimunlidae
Harrimania
kupJferi (von Willemoes-Suhm, 1 87 1 )
SaccogJossus
horsl; BrambeH & Goodhart, 1 9 4 1
kowalevskii (Agassiz, 1 873)
mereschkawskii (Wagner, 1 885)
pygmaeus Hinrichs & Jacobi, 1 93 8
ruber Tattersall, 1 905
Stereobalanus
canadensis (Spengel, 1 893)
Prologlossus
kaehleri (Caullery & Mesnil, 1900)
Family Ptychoderidae
Balanoglossus
clavigerus delle Chiaje, 1 829
Glossobalanus
elongatus Spengel, 1904
marginalus Meek, 1 922
minutus Kowalevsky, 1 866
sarniensis (Koehler, 1 886)
Family SpengeJidae
Glandiceps
ta/aboti (Marion, 1 876)
Family il/cer/ae sedls
Planctosphaera
pelagica (Spengel, 1 932)
Class Pterobrancbia
Family Rhabdopleuridae
Rhabdopleura
compacta Hincks, 1 880
normani Allmann, 1 869
ECHINODERMATA
Compiled by Hans G. Hansson
The list of European Echinodermata was compiled by
Hans G. Hansson. H used the Echinoderm part of the
compilers NEAT lists (available via Internet at address
htttp:llwww.tmbl.gu .se/li bdb/taxon/taxa.htm l) as a
starting point. NEAT (North East Atlantic Taxa) are
preliminary check-lists, covering Scandinavian marine
animal taxa. The chapter by FJ. Fell on Echinodermata
in Parker ( 1 982) was used as the systematic framework.
A The asteroids are mainly listed from the recent
monograph on Atlantic species by Clark & Downey
A ( 1 992). Mortensen ( 1 927) is used as a basal source for
A
the other groups. Additions from several other sources
AM
A
have been made, and the electronic part of the Zoological
A
Record (covering recent decades) has been utilised.
However, examination of the reports from the large
A expeditions (except the Norwegian North Atlantic and
the (ngolf) and other literature since WWII, may add to
A the species list.
The compiler is generally interested in marine fauna, but
is not a specialist on any Echinoderm taxon, so there are
AM
certainly several omissions and lapses i n this draft list.
M The assistance of experts in groups of Echinodermata
A was thus invaluable. Dr Sabine StOhr, Stockholm,
AM assisted w ith ophiuroid names, Dr Claude Massin,
A Brussels, assisted with the holothurians and added 9
omitted species of this taxon to the list. Dr Andrey
Gebruk, Moscow, together with his friends Alexander
M Mironov and Alexei Smirnov kindly helped with several
corrections and additions to the holothurians, the
crinoids, the echinoids and the genus Caulaster among
A the asteroids. Prof. Dusan Zavodnik, Rovinj, Croatia,
provided severa l corrections, new distribution notes
(particularly for the Adriatic Sea area), information about
synonyms, and added a few omitted species. Dr Mateo
A Garrido, Gran Canada, provided additional records from
AM the Canaries.
Authors of most genus and species names in the list
below have been found, and most often also the years of
description, but with several exceptions. Also, some of
the years of authority in this list may be wrong, because
different sources often have different opinions regarding
which year a certain species is described (e.g. gen.
Poliometra Clark; the original publication: 1 923, Neave:
1 924, McKenzie in Howson & Picton 1 997: 1 9 1 3; in this
case the year from the original publ ication must be
correct). Thus, further study of the primary literature
may improve the accuracy of this list.
References
Belyaev G.M. & A.N. Mironov, 1 996. Starfishes Porcellanaster
and Caulaster (Porcellanasteridae) from the Atlantic and
the Antarctic. Zoologichesky zhurnal, 75 (6): 886 - 899 (In
Russian, English summary).
Brattegard, T & T. Holthe (Eds.). 1 997. Distribution o/marine.
benthic macro-organisms in Norway. A tabulated
catalogue. Preliminary edition. Reseach Report for ON, Nr.
1 997- 1 . Trondheim. 409 pp. (Echinoderm chapters by T.
HBis8!ter, T Brattegard & J.-A. Sneli, pp. 3 0 1 -3 1 5).
Clark, A.H. 1 923. Crinoidea. The Danish Ingolf Expedition . .
Vol. IV. Luno, Copenhagen. 5 8 pp.
Clark, A.M. 1 970. Echinodermata Crinoidea. Marine
Invertebrates a/Scandinavia. 3. Universitetsforiaget, Oslo.
55 pp.
Clark, A.M. & M.E. Downey 1 992. Starfishes o/the Atlantic.
Chapman & Hall, London. 794 pp.
- 3 36 -

D'yakonov, A.M. 1 967. Ophiuroids o/the USSR Seas. Israel
Program for Scientific Translations, Jerusalem 123 pp.
D'yakonov, A.M. 1 968. Sea Stars (Asteroids) o/the USSR Seas.
Israel Program for Scientific Translations, Jerusalem 1 83
pp.
D'yakonov, A.M. 1 969. Echinodermata, Vol. 1. Sea-Urchins
(Echinoidea) No. I. Fauna 0/Russia and adjacent
Countries. Israel Program for Scientific Translations,
Jerusalem 265 pp.
Fell, H.B. 1 960. Synoptic Keys to the Genera of Ophiuroidea.
Zoology Publicalionsfrom Victoria University of
Wellington No. 26. 34 pp + literature index.
Fiege, D. & Y. Liao 1 996. Penilpidia IUdwigi (Marenzeller,
1 893) (Holothurioidea: Elpidiidae) rediscovered from the
Eastern Mediterranean Sea. In: Uiblein, F., OIt,I.,
Stachowitsch, M. (Eds) Deep-sea and extreme shallow
water habitats: affinities and adaptations. Biosystematics
and Ecology Series, 1 1 , 61 -66.
Gebruk, A.V. 1 989. Revision of the family Pelagothuriidae
(Holothurioidea, Elasipodida) with a review of swimming
holothurians. Part I . Revision of Pelagothuriidae. Zool.
Zhurnal 68 ( 1 2): 57-66. (In Russian, English summary).
Gebruk, A.V., 1 990. Deep-sea holothurians ofthe/amily
Elpidiidae. Moscow, Nauka, 1 60 p. [In Russian].
Gulliksen , B. Palerud, R., Brauegard, T. & I.-A. Sneli, 1 999.
Distribution of marine benthic macro-organisms at
Svalbard (including Bear Island) and Jan Mayen. Research
report for DN 1 999 - 4. 1 4 8 pp. Trondheim, Norway.
Hansen, B, 1 975. Systematics and biology of the deep-sea
holothurians. Galathea Report, 13, 1 -262.
Heding, S.G. 1 935. Holothurioidea I. The Danish Ingolf
Expedition. Vol. IV. Hagerup, Copenbagen. 84 pp + 8
plates.
Heding, S.G. 1 93 5 Echinoderms. Meddele/ser om Gren/and.
Bd. 1 04, Nr. 1 3. 68 pp + 1 plate.
Heding, S.G. 1 936 Echinoderms. Meddelelser om Gren/and.
Bd. 1 08, Nr. I . 34 pp.
Heding, S.G. 1 942. Hololhurioidea II. The Danish Ingolf
Expedition. Vol. IV. Hagerup, Copenbagen. 39 pp + 2
plates.
Heding, S.G. & A. Panning 1 954. Phyllophoridae. Spolia
Zoologica Musei Hauniensis XIII. 209 pp.
Howson, C.M. & B.E. Picton 1 997 (Eds.). The Species
Directory o/the Fauna and Flora o/the British Isles and
Surrounding Seas. The Ulster Museum, Belfast & The
Marine Conservation Society, Ross-on-Wye. 508 pp.
(Echinoderm chapter by J.D. McKenzie, pp. 287-295).
Heisa:ter, T. 1 990. An annotated check-list of the echinoderms
of the Norwegian coast and adjacent waters. Sarsia 75 : 83-
1 06
Kier P.M. & M. H. Lawson, 1 978. Index of Living and Fossil
Echinoids 1 924- 1 970. Smithson. Contrib. Paleobiol. 34 :
vi+ 1 82 pp. (not yet seen)
Madsen, FJ. 1 947. The Echinoderms collected by the
"Skagerak" expedition in the eastern Atlantic 1 946. I.
Asteroidea, Ophiuroidea, Echinoidea, and Holothuroidea.
Goteborgs Kungl. Vetenskaps- och Villerhets-Samhiilles
Handlingar, Sj !ltte fOljden. Ser. B, Band 5. N:o 7. 1 6 pp.
Madsen, F.J. 1 987. The Henricia sanguino/enta complex
(Echinodermata, Asteroidea) of the Norwegian Sea and
adjacent waters. A re-evaluation, with notes on related
species. Steenstrupia 1 3 (5), pp. 20 1-268.
Madsen, FJ. & B. Hansen 1 994. Echinodermata
Holothurioidea. Marine Invertebrates of Scandinavia. 9.
Scandinavian University Press. 1 43 pp.
M i l ler, J.E and Pawson, D.L. 1 990. Swimming sea cucumbers
(Echinoderamata :Holothuroidea): a survey, with analysis of
swimming behaviour in four bathyal species. Smithsonian
Contributions to Marine Sciences, 35, 1 8p.
Mironov A.N., 1 974. Pourtalesiid sea urchins ofthe Antarctic
and Subantarctic (Echinoidea: Pourtalesiidae). Transactions
o/the P.P. Shirshov Institute 0/oceanology, 98: 240 - 252.
(In Russian, English summary).
Mironov A.N., 1 985. Circumcontinental zonation in distribution
of Atlantic echinoids. Transactions o/the P.P. Shirshov
- 337 -
European Register of Marine Species
Institute 0/oceanology, 1 20, 70-95. (In Russian, English
summary).
Mironov A.N. 1 995. Holasteroid echinoids. 2. Pourtalesia.
Zoologichesky zhurnal, 74 ( 1 2), 59-75. (In Russian, English
summary).
Mironov A.N. 1 997. Holasteroid echinoids. 4. Echinosigra.
Zoologichesky zhurna/, 76, (2): 1 73-1 86. (In Russian,
English summary).
Mortensen, T. 1 903. Echinoidea I. The Danish lngolf
Expedition. Vol. IVa. Luno, Copenhagen. 1 83 pp + 2 1
plates.
Mortensen, T. 1 907. Echinoidea II. The Danish Ingolf
Expedition. Vol. IVa. Luno, Copenhagen. 200 pp + 1 9
plates.
Mortensen, T. 1 927, Handbook ofthe Echinoderms 0/the
British Isles. Clarendon Press, Oxford. 47 1 pp.
Mortensen, T. 1 933. Ophiuroidea. The Danish lngolf
Expedition. Vol. IV. Luno, Copenhagen. 1 2 1. pp + 3 plates.
Neave, S.A. 1 939-50. Nomenclator Zoologicus . Vol . l 5 . The
Zoological Society of London. + M.A. Edwards & A.T.
Hopwood (Eds.) 1 966 Vol. 6 + M.A. Edwards & H.G.
Vevers (Eds.) Vol. 7 .
Parker, S.P. (Ed.) 1 982. Synopsis and Classification 0/ Living
Organisms. 2 Vols. McGraw-Hili, New York. 1 166 + 1 232
pp.
Paterson, G .L.J. 1 985. The deep-sea Ophiuroldea of the north
Atlantic Ocean. Bulletin o/the British Museum (Natural
History). Zoology. 49 ( I ). 1 62 pp. (not seen)
Pereira, Mariano Oliveiro Rego 1 997. Checklist of the l ittoral
echinoderms of the Azores. Aforeana 8 (3): 3 3 1 -337.
Picton, B.E. 1 993. A Field Guide to the Shallow-water
Echinoderms o/the British Isles. Immel Publishing. 96 pp.
Sherborn, C.D. 1 902. Index Animalium sive index nom inurn
quae ab a.d MDCCL VIII generibus et speciebus anlmalium
imposita sunt. Sectio prima a kalendis Ianuariis,
MDCCLVIII usque adjinem Decembris. MDCCC. C. J.
C lay & Sons, Cambridge Unlv. Press, London. 1 1 95 pp.
Sherborn, C.D. 1 922-32. Index Animalium sive index nominum
quae ab a,d MDCCLVIIl generibus et speciebus animalium
imposita sunt, Sectio secunda a kalendis Ianuariis,
MDCCCI usque adjinem Decembris, MDCCCL. The
Trustees of the British Museum, London. 7056 + 1098 pp.
Tortonese, E. 1 965. Echinodermata. Fauna d'italia. Vol. 6.
Edizioni Calderini, Bologna. 422 pp.
PHYLUM ECIDNODERMATA
Class Crinoidea
Order Millericrinida
Family Hyocrinidae
Gephyrocrinus
grima/dii Koehler & Bather, 1 902
Order Cyrtocrinida
Family Holopodidac
Cyathidium
/oresti Cherbonnier & Guille, 1 972
C. meteorensis Fechter, 1 973
Order Bourgueticrinida
Family Bourgueticrinidae
Conocrinus
cabioshi Roux, 1 976
cherbonnieri Roux, 1 976
Democrinus
par/aUi Perrier, 1 883
Rhizocrinus magnus Gislen, 1 947
=
Rhizocrinus rawsoni Koehler, 1 909, non Pourtales,
=
1 874
conifer (A.H. C lark, 1909)
Bythocrinus intermedius A.H. Clark, 1 9 1 5
Rhizocrinus conifer A.H. Clark, 1 909
Rhizocrinus robustus A.H. Clark, 1 909
=
rawsonii (Pourtali!:s, 1 874)
Rhizocrinus rawsonii Pourtales, 1 874
=
Rhizocrinus
Costello, Emblow and White (eds), 200 1

lofotensis M. Sars, 1 868 Trichometra

= R. verrilli A.H. Clark, 1 908 cubensis (Pourtales, 1 869)
Family Bathycrinidae Antedon eubensis Pourtales, 1 869
=

Bathycrinus T. aspera AH. Clark, 1 908

aldrichianus Wyville Thomson, 1 817 delieata A.H. Clark, 1 9 1 J
= B. serratus AH. Clark, 1 908 Thaumatomelra
carpenlerii (Danielssen & Koren, L 811) seplentrionalis A.H. Clark, 1 923
lIycrintls carpenferii Danielssen & Koren, 1 811 Family Atelecrinidae
gracilis Wyville Thomson, 1 817 A telecrinus
Monachocrinus balanoides P.H. Carpenter, 1 88 1
perrieri (Koehler & Vaney, 1 9 1 0) =A. helgae A.H. Clark, 1 9 1 3
= Balhycrintls perrieri Koehler & Vaney, 1 9 1 0 Family Comasteridae
M coe/us H.L. Clark, 1 923 Neocomalella
recuperattls (Perrier, L 885) europaea A.H. Clark, 1 9??
= Ilycrinus rectlperalus Perrier, 1 885
Family Thalassometridae
sexradiatus AH. Clark, 1 9 1 9
Koehlermelra
Family Phrynocrinidae f1ava (Koehler)
Zeuclocrinus =Anledonflava Koehler
gis/eni AM. Clark, 1 913 porrecta (P.H. Carpenter, 1 8??)
Family Porphyrocrinidae =Antedon porrecla P.H. Carpenter, 1 8??
Porphyrocrinus Thalassometra
incrassalus (Gislen, 1933) lusilanica (P.H. Carpenter, 1 884) A.M. Clark, 1 980
Ihalassae Roux, 1 911 =Antedon lusilanica P.H. Carpenter, 1 884
Sliremelra ltlsitaniea (P.H. Carpenter, 1 884)
Order Isocrinida
omissa (Koehler)
Family Isocri nidae =Antedon omissa Koehler
Diplocrinus
(Annacrinus) wyvillelhomsoni (Jeffreys, 1 8??) Class Asteroidea
Penlacrinus wyville-Ihomsoni Jeffreys, 1 8?? Order Paxillosida
Order Comatulida Family Aslropectinidae
Family Pentamctrocrinidae Asrropecten
Penlamelrocrinus arandacus (Linnaeus, 1 158)
allanticus (Perrier, 1 883) Aslerias arandaca Linnaeus, 1158
=

Eudioerinus allanticus Perrier, 1 883 Aslerias auranliaca Tiedemann, 1 8 1 6

=

Thaumalacrinus A. antarcticus Studer, 1 816

jungerseni A.H. Clark, 1 923 A. crenaster Dujardin & Hupe, 1 862
=

A. gruveN Koehler, 1 9 1 2
Family Antedonidae
A . meridionalis Studer, 1 876
=

Antedon A. perarmalus Perrier, 1 869

=

bifida (Pennant, 1 117) bispinosus (Otto, 1 823)

?Anledan duebeni Bohlsche, 1 866
Aster/as bispinosa Otto, 1 823
=
=
A. gorgonia de Freminville, 1 8 1 1
A. myosurus Perrier, 1 869
=

A. maroccana A.H. Clark, 1 9 1 0 hermalophilus Siaden, 1 883

Aslerias bifida Pennant, 1 711
iberieus Perrier, 1 894
=

medilerranea (de Lamarck, 1 8 1 6) irregularis subsp. irregularis (Pennant, 1 771)

= A. adriatica A.H. Clark, 1 9 1 1 Asterias irregularis Pennant, 1 171
=

Comalula annulala Risso, 1 826

A. acleularis Nonnan, 1 865
=
=

Comalula medilerranea de Lamarck, 1 8 1 6 A. echinulatus J. MUller & Troschel, 1 844

=

pelasus (Dahen & Koren, 1 846) A. helgolandieus Greeff, 1 87 1

=

Aleelo petasus DUben & Koren, 1 846

A. muelleri }. MUlier & Troschel, 1 844
=
=

Hathrometra Irregu/ris subsp. pentaeanlhus (Delle Chiaje, 1 827)

sarsii (DUben & Koren, 1 846) Aster/as pentacantha Delle Chiaje, 1 827
=

Antedon teneila Carpenter non (Retzius)

jonsloni (Delle C hiaje, 1 827)
= H. tenel/a (Carpenter, 1 888) non (Retzius, 1 783) Asterias jonstoni Delle Chiaje, 1 827
=

Heliometra A. aster De Filippi, 1 859

=

glaeiaUs (Owen, 1 83 3 ex Leach MS)

A. squamalus J . MOller & Troschel, 1 844
=

Aleelo eschrichtii J. MUller, 1 84 1

platyacanlhus (Philippi, 1 837)
=

= Alecto glacialis Owen, 1 83 3 ex Leach MS Aslerias platyacantha Philippi, 1 837

=

Antedon quadrata P .H. Carpenter, 1 884

spinulosus (Philippi, 1 831)
=

Leptometra Asterias spinulosa Phil ippi, 1 837

=

cellica (M'Andrew & Barrett, 1 858) Bathybiasler

Comalula cel1ica M'Andrew & B arrett, 1 85 8
vexilfifer (W. Thomson, 1 873)
=

= Comatula woodwardi Barrett, 1 857, non Forbes, 1 852 Archaster robuslus Verrill, 1 884
=

L. phalangium (Carpenter, 1 8 8 1 ) non (J. MUller, 1 84 1 )

Archasler vexillifer W. Thomson, 1 873
=

phalangium (J. MUlier, 1 84 1 ) Aslropeelen pallidus Danielssen & Koren, 1 817

=

= Aleclo phalal1gium J. MUlier, 1 8 4 1 =Ifyasler mirabilis Danielssen & Koren, 1 884

Meteororomelra Phoxaster pumilus Siaden, 1 889
montieola A.M. Clark, 1 980 Dytasler
Orthometra grandis subsp. grandis (Verrill, 1 884)
hibernica (A.H. Clark, 1 9 1 3) Archasler grandis Verrill, 1 884
= Trichometra hibernica A.H. Clark, 1 9 1 3 =D. agassizi Perrier, 1 894
Poliometra D. biserialis Siaden, 1 889
prolixa (Sladen, 1 88 l ) =D. exilis var. carinala S laden, 1 889
= Anledon prolixa Siaden, 1 88 1 =D. madreporifer Sladen, 1 889
Harhrometra prolixa (Sladen)

- 338 -
 European Register of Marine Species

D. rigidus Perrier, 1 894 = H. fortis Koehler, 1 907

intermedius Perrier, 1 89 1 = H. gracilis Koehler, 1 907
moliis (perrier, 1 885) = H. parfaiti Perrier, 1 885
Crenaster mollis Perrier, 1 885 Parcel/anaster
semispinosus (perrier, 1 894) ceruleus Wyville Thomson, 1 87 7
Crenaster semispinosus Perrier, 1 894 Albatrossia semimarginalis Ludwig, 1 905
=

D. parvulus Koehler, 1 909 Cau/aster sladeni Perrier, 1 885

Leplychaster P. granulosus Perrier, 1 88 5
=

arctjcus (M. Sars, 1 85 1 ) P. inermis Perrier, 1 885

=

Astropecten arct/cus M . Sars, 1 85 1 P. irreglilaris Lieberkind, 1 932

As/ropec/en lutken; Barrett, 1 857 Slyracasler
Loncho/as/er armalus Siaden, 1 883
/ar/areus Sladen, 1 889 Machairaster pictus Perrier, 1 884
=

Persephonaster S. edwardsi Perrier, 1 885

=

/eptactis H.L. Clark, 1 94 1 S. spinosus Perrier, 1 885

patagia/us (Sladen, 1 889) chuni Ludwig, 1 907
P. spinulosus H.L. Clark, 1 94 1 elongallis Koehler, 1 907
Psi/aster patagia/us Siaden, 1 889 horridus Siaden, 1 883
sphenoplax (Bell, 1 892) Chunasler scapanephorus Ludwig, 1 907
=

Astropec/en sphenop/ax Bell, 1 892 Thoracas/er

= Psi/asteropsis humilis Koehler, 1 907 eylindrailis Sladen, 1 883
P/ulonaster Pseudaster cordi/er Perrier, 1 885
=

agassizi subsp. nota/us S iaden, 1 889 T. magnus Ludwig, 1 907

= ?Goniopecten edwardsi Perrier, in Milne-Edwards, Order Notornyotida
1 882
?Goniopecten inermis Perrier, 1 885 (nomen nudum) Family Benthopectinidaf
= P. abbreviatus Siaden, 1 889 Benthopeelen
P. granulosus Perrier, 1 89 1 simplex subsp. simplex (Perrier, 1 88 1 )
= P. no/alus S laden, 1 889 Archaster simp/ex Perrier, 1 88 1
bifrons (Wyvil\e Thomson, 1 873) B. spinasus Grieg, 1 9 1 8 (p.p.), non Verrill, 1 884
=

Archas/er bifrons Wyville Thomson, 1 873 = Pararchas/er armatus Siaden, 1 889

= P. proteus H.L. Clark, 1 92 3 spinosissimus (Sladen, 1 889)
Psi/aster = Pararchas/er spinosissimus Siaden, 1 889
andromeda subsp. andromeda (J. MUlier &. Troschel, Cheiraster
1 842) sepilus (Verrill, 1 885)
Astropecten andromeda J. MUller &. Trosche1, 1 842 = Archaster sepi/us Verrill, 1 885
Astropecten crisUi DUben &. Koren, in DUben, 1 845 Luidiaster dubius H.L. Clark, 1 94 1
cassiope Sladen, 1 889 =Peclinaster dispar Verrill, 1 9 1 5
?Archaster pulcher Perrier, 1 88 1 =Pectinaster echinu/atus Ludwig, 1 9 1 0 (p.p.)
= P. andromeda var. cassiope Grieg, 1 907 Pontaster perp/exlls Perrier, 1 894
= P. squameus H.L. Clark, 1 94 1 =Pontaster venus/us Siaden, 1 889
Tethyaster Peetinas/er
subinermis (Philippi, 1 837) fllholi Perrier, 1 885
Asterias subinermis Philippi, 1 837 =Pan/aster forcipatus Siaden, 1 889
Pantaster prislinus Siaden, 1 889
Family Luidiidae
Ponlaster
Luidia lenuispinus (DUben &. Koren, 1 846)
atlanlidea Madsen, 1 950 =Astropeelen lenuispinus DUben &. Koren, 1 846
ciliaris (Philippi, 1 837) =Crenaster marionis Perrier, 1 885
Aslerias imperali Delle Chiaje, 1 84 1 =P. hebilus Siaden, 1 889
= L. fragilissima Forbes, 1 839 P. limbatus Siaden, 1 889
sarsil subsp. sarsii D!lben &. Koren, in Dilben, 1 845
Astrella simp/ex Perrier, in Mi lne-Edwards, 1 882 Order Va)vatida
= L. africana S iaden, 1 889 Family Chaetasteridae
FamilY Ctenodiscidae Chaetaster
Ctenodiscus /ongipes (Relzius, 1 805)
crispatus (Relzius, 1 805) Asterias longipes Retzius, 1 805
=

Asterias aranda Dewhurst, 1 834 Asterias subu/ata de Lamarck, 1 8 1 6

=

= Asterias crispala Retzius, 1 805 Nepanlhia /esselala Gray, 1 840

= Asterias palaris Sabine, 1 824 Family Odontasteridae
C. cornicu/atus Perrier, 1 875 Hop/asler
= C. pygmaeus J. MUlier &. Troschel, 1 842 spinosus Perrier, in Milne-Edwards, 1 882
Family Radiasteridae = Penlagorwsler /epidus Siaden, 1 889
Radiaster Odontaster
lizardi (Sladen, 1 882) mediterraneus (von Marenzeller, 1 893)
Mimasler lizardi Sladen, 1 882 Gnathasler medilerraneus von Marenzeller, 1 893
=

Family Porcellanasteridae Family Goniasleridae

Cau/aster Ceramasler
eremicus (Sladen, 1 889) granularis subsp. granularis (Relzius, 1 783)
= A/balrotrossaster richardi Koehler, 1 909 Aslerias granularis Retziu8, 1 783
=

Porcel/anaster eremicus Siaden, 1 889 Aslerias tessellatus de Lamarck, 1 8 1 6

=

pedlinculailis Perrier, 1 882 Astrogonium borea/e B arrett, 1 857

=

Hypha/asler = Pentagonaster eximius Verrill, 1 894

inermis Siaden, 1 883 Pentagonaster simp/ex Verrill, 1 895
= H. anlonU Perrier, 1 885 grenadensis subsp. grenadensis (Perrier, 1 88 1 )

- 339 -
Costello, Emblow and White (eds), 2001

= Astrogonium greeni Bell, 1 889 Aphroditasler gracilis Siaden, 1 889

=

= Penlagonaster qffinis Perrier, 1 884 ASlrogonium aequabile Koehler, 1 907

=

= Penlagonasler balteatus Siaden, 1 8 9 1 Aslrogonium aphrodite Perrier, 1 894

= Pentagonaster crassus Perrier, 1 885 Astrogonium eminew Koehler, 1 907
=

= Pentagonaster deplasi Perrier, 1 88 5 ASlrogonium marginalum Koehler, 1 909

= Pentagonaster gosselini Perrier, 1 884 Astrogonium necator Perrier, 1 894
= Penlagonaster grenadensis Perrier, 1 8 8 1 P. cancinnus Verril l , 1 894
=

Pentagonaster haesitans Perrier, 1 885 P. granuliferus Verrill, 1899

= Penlagonaster hyslricis von MarenzeIler, 1 893 P. ordina/us Verrill, 1 899
=

= Penlagonaster kergroheni Koehler, 1 895 parelii (D!lben & Koren, 1 846)

Evoplosoma Aslrogonium annectens Perrier, 1 894
scorpio Downey, 198 1 = Astrogonium/allax Perrier, 1 885
Goniasler As/rogonium hystrix Perrier, 1 894
tessellatus (de Lamarck, 1 8 1 6) Aslrogonium longobrachia/e Koehler, 1 907
= Asterias granufaris Gmelin, 1 79 1 (p.p.) Astropecfen parelii Dilben & Koren, 1 846
=

Aslerias lesselfata de Lamarck, 1 8 1 6 P. intermedius var. insignis Verrill, 1 89 5

= Astrogonium duhium Perrier, 1 869 = P. intermedius S iaden, 1 889
= G. africanus Verrill, 1 87 1 = P. tessel/alus var. arcticus Sluiter, 1 895
= G. americanus Verrill, 1 87 1 Sphaeriodiscus
= G. cuspidafus Gray, 1 840 bourgeli (Perrier, 1 885)
G. sehae Gray, 1 840 = Slephanasier bourgeti Perrier, 1 885
Hippasferia Family Ophidiasteridlle
phrygiana subsp. phrygiana (Parelius, 1 768) Hazelia
Aslerias equestris de Lamarck, 1 8 1 6, non Retzius, 1 805 a/fenuala Gray, 1840
= Asterias johnstoni Gray, in Johnston, 1 836 Aslerias coriacea Grube, 1 840
=

= Asterias phrygianus Parelius, 1 76 8 Asterias laevigala subsp. varielas de Lamarck, 1 8 1 6

=

Astrogonium acufeatum Barrett, 1 857 = ASlerias variolala Risso, 1 826, non Retzius, 1 805
= Goniaster ahbensis Gray, 1 843 Ophidiasler (Hazelia) atfenualus Gray, 1 84 0
=

H. (Nehippasleria) insignis Dons, 1 93 8 Ophidiaster lessonae Gasco, 1 876

= H. cornula Gray, 1 840 Narcissia
= H. europaea Gray, 1 84 0 canariensis (d'Orbigny, 1 839)
H. plana Gray, 1 840 A slerias canariensis d'Orbigny, 1 839
=

insignis Dons, 1 937 Fromia narcissae Perrier, 1 885

=

Mediasler Fromia narcissiae Perrier, 1 894

bairdi subsp. bairdi (Verrill, 1 882) N. lenerifJae Gray, 1 840
= Archasler bairdi Verrill, 1 882 Irigonaria Sladen, 1 889
= M slellatas Perrier, 1 89 1 Ophidiaster
Nymphaster ophidianus (de Lamarck, 1 8 1 6)
arenalus (Perrier, 1 8 8 1 ) Asterias ophidiana de Lamarck, 1 8 1 6
=

Dorigonajacqueli Perrier, 1 894 O. aurantius Gray, 1 84 0

= Dorigona prehensilis Perrier, 1 88 5 O. canariensis Greeff, 1 872
=

= N. albidus S laden, 1 889 reyssi Sibuet, 1 977

N. basilicus S laden, 1 88 9
Family POfaniidae
= N. prolenlus Siaden, 1 889
Chandraster
= Penlagonasler arena/as Perrier, 1 88 1
grandis (Verrill, 1 878)
= Penlagonasler subspinosus Perrier, 1 88 1
?Marginasler fimbrialus Siaden, 1 889
= Penlagonas/er lernalis Perrier, 1 8 8 1
=

=Porania grandis Verrill, 1 878

Paragonaster
Marginasler
subliUs (Perrier, 1 88 1 )
capreensis (Gasco, 1 876)
=Archasler/ormosus Verrill, 1 884
Asterops/s capreensis Gasco, 1 876
Goniopeclen sublilis Perrier, 1 88 1
=

Porania
= P. cylindralus Siaden, 1 889
(Porania) pu/villus subsp. pulvillus (O.F. Maller,
= P. strictus Perrier, 1 894
1 776)
Pentagonaster elongatus Perrier, 1 88 5
Asleropsis clenacanlila J. Maller & Troschel, 1 84 2
=
Pellasler
= Ganiaster templeloni Thomson, in Forbes, 1 839
placenla (J. MUller & Troschel, 1 842)
P. gibbosa Gray, 1 840 (nomen nudum)
Gonias/er nidarosiensis Storm, 1 8 8 1
(Pseudaporania) starmi (Dons, 1 935)
= Goniodiscus aculus Heller, 1 863
Pseudaparania stormi Dons
Goniodiscus placenla 1. MUller & Troschel, 1 842
=

Poraniomorpha
= Goniodiscus placenlae/ormis Heller, 1 863
(Poraniomorpha) bidens Mortensen, 1 93 2
= P. hebes Verrill, 1 899
(Poraniomorpha) hispida subsp. hispida (M. Sars,
Penlaganasler minor Koehler, 1 895
1 872)
= Penlagonasler mirabilill Perrier, 1 875
= ?Rhegasler murrayi Sladen, 1 883
= Penlagonasler planus Verrill, 1 895
=Asterina borealis Verrill, 1 878
= Penlagonasler vincenti Perrier, 1 88 5
=Goniaster hispidus M. Sars, 1 872
PUnlhasler
= Parania spinulosa Verrill, 1 880
denialUs (Perrier, 1 884)
(Poraniamorpha) hispida subsp. rosea Danielssen &
Penlagonaster concinnus Siaden, 1 89 1
Koren, 1 8 8 1
= Pentagonasler denlalus Perrier, 1 884
= Lasiaster villosus Siaden, 1 889
Pentagonasler grandis Perrier, 1 88 5
P. rosea Danielssen & Koren, 1 8 8 1
= Pentagonasler perrieri Siaden, 1 889
=

(Poraniomorpha) tumida (Stux.berg, 1 878)

Tosia (PUn/haster) compla Verrill, 1 899
Rhegasler lumidus (Stuxberg) auctt.
Tosia (Plinthaster) nilida Verrill, 1 899
=So/asler tumida Stux.berg, 1 878
Pseudarchaster
=Sphaeriasler bjoerklykkei (Dons, 1 938)
gracilis subsp. gracilis (Sladen, 1 889)
(Culci/opsis) borealis (SOssbach & Breckner, 1 9 1 1 )

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 European Register of Marine Species

?Marginaster perllagonlls Perrier, in Milne-Edwards, = H. knipowitschi D'yakonov, 1950

1882 Ii. sanguinolenta Rasmussen, 1965 non (O.F. Mllller)
CII/cila borealis SOssbach & Breckner, 1911 pertusa (O.F. Mllller, 1776)
Sphaeraster berthae Dons, 1938 ?Cribrefla biseayensis Koehler, 1895
= Sphaer/aster berthae Dons, 1939 = Asterias pertllsa O.F. MOiler, 1776
Ty/aster = Cribrella sanguinoienta var. abyss/cola Norman, 1869
wille; Danielssen & Koren, 1881 Eehinaster oelllatus f. major Dilben & Koren, 1846
Family Sphaerasterldae sanguinolenta (O.F. Mllller, 1776)
Podosphaeraster = Asterias sanguinolenta O.F. MOiler, 1776
tha/assae Cherbonnier, 1970 Cribrella pectinata Verrill, 1894
= Cribrella sanguino/enta var. curta Norman, 1869
Family Asterlnidae
Cribrella sanguinolenta f. lawior Michailovskij, 1902
A lIopatiria = Echinaster oculatlls f. minor Dilben & Koren, 1846
ocelllfera (Gray, 1847) = Echinaster sarsi J. MUlier & Troschel, 1844
= ?Palmipes lobiancoi Ludwig, 1897 Ii. pCI/orata Rasmussen, 1965 non (O.F. MUller)
Parasterina africana Engel & eroes, 1960 =Ii. sanguinoienla subsp. miliaris Verrill, 1914
Paliria ocelllfera Gray, 1847
skorikovi D'yakonov, 1950
= Patiria pulla Koehler & Vaney, 1906 solido D'yakonov, 1950
= Patiria rosea Koehler & Vaney, 1906 spongiosa (0. Fabricius, 1780)
Anseropoda
lobiancoi (Ludwig, 1897) Order Velatida
Palmipes lobiancoi Ludwig, 1897 Family Soilisteridae
placenta (Pennant, 1777) Crossaster
= Asterias carti/aginea Fleming, 1828 papposus (Linnaeus, 1767)
= Aster/as membranacea Retzius, 1783 Asterias qffinis Brandt, 1835
= Aster/as placenta Pennant, 1777 = Asterias helianthemoides Pennant, 1777
= Aster/scus palmipes J. MOller & Troschel, 1842 = Asterias papposa Linnaeus, 1767
Asterina sqllamallls (Dllderlein, 1900)
gibbosa (Pennant, 1777) = So/asler affinis Danielssen & Koren, 1877 non (Brandt,
Asterias gibbosa Pennant, 1777 1835)
= Asterias verruculata Retzius, 1805 = SO/OSler papposus var. squamata D1lderlein
= A. minllta Nardo, 1834 = So/asler squamatus D6derlein
= AsterisellS arrecijiensis Greeff, 1872 Lophaster
pancerli (Gasco, 1870) /urelfer (DUben & Koren, 1846)
= AsterisellS paneerii Gasca, 1870 = Solaster /urclfer Dilben & Koren, 1846
phylaeliea Emson & Crump, 1979 Solaster
wega Perrier ear/li Verrill, 1879
Tremaster endeca (Linnaeus, 1771)
mirabilis Verrill, 1880 Aster/as alboverrueosa Brandt, 1835
T. lawis H.L. Clark, 1941 = Asterias aspera O.F. MOiler, 1776
Order Spinulollida = Asterias endeea Linnaeus, 1771
= Asterias rumphii Parelius, 1768
Family Echinasteridllc
= S. endeca subsp. deeemradiata Siaden, 1889
Echinaster S. galaxides Verrill, 1909
(Eehinaster) sepositus subsp. sepositus (Retzius, = S. intermedius Sluiter, 1895
1783) glaeialis Danielssen & Koren, 1881
= Asterias sagena Retzius, 1805 = S. echinatus Storm, 1888
= Asler/as seposita Relzius, 1783 syrtensis Verrill, 1894
= Rhopia mediterranea Gray, 1840
Family Korethrasteridae
Henricia
Korethrasler
caudani (Koehler, 1895)
h/spidlls Wyville Thomson, 1873
= Cribella caudani Koehler, 1895
K. selosus Perrier, in Milne-Edwards, 1882
cylindrel/a (Sladen, 1883)
= Cribrella abyssa/is Perrier, 1894 Family Plerasteridae
= Cribrella oculala var. eylindrella Siaden, 1883 Diplopteraster
H. abyssicola Perrier, 1885 & Farran, 1913 & multipes (M. Sars, 1866)
Mortensen, 1927 = Pterasler mllil/pes M. Sars, 1866
esehrichti (J. MOiler & Troschel, 1842) Hymenaster
= Eehinaster esehriehti J. MUller & Trosehel, 1842 giboryi Perrier, 1894
= Magda/enaster arct/cus Koehler, 1907 pellucidus Thomson, 1873
hedingi Madsen, 1987 = Ii. giganteus Siaden, 1891
lisa A.H. Clark, 1949 = H. membranaceus Thomson, 1877
lisa subsp. ingolfl Madsen, 1987 = H. perissonotus Fisher, 1910
ocu/ata (pennant, 1777) regalis Verrill, 1895
= Asterias oeulata Pennant, 1777 = H. kieri Downey, 1979
Ii. sanguinolenta auelt., non (O.F. MOiler) (p.p.) reticulatus Sibuet, 1976
per/orata (O.F. MUller, 1776) rex Perrier, 1885
= Asterias per/orata O.F. MOiler, 1776 = H. gennaeus H.L. Clark, 1923
= Cribrefla oeulata var. murina Perrier, 1875 rosells Koehler, 1907
= Cribrefla sanguino/enla f. seabrior Michailovskij, 1902 tenllispinlls Sibuet, 1976
= Echinaster serobieulalus Danielssen & Koren, 1883 Pleraster
Ii. sanguino/enta auett., non (O.F. Muller) ac/cllia (Downey, 1970)
= Ii. cross/spika Heding, in Jltgerski61d, 1971 (nomen = Marsipasler acicula Downey, 1970
nudum) a/veolaills Perrier, 1894
= Ii. fisheri subsp. echinala Heding, in J!lgerski6Id, 1971 hystrix Harvey, 1989
(nomen nudum) mifital'is (O.F. MUlier, 1776)

- 341 -
Costello, Emblow and White (eds), 200 I

= Aslerias militaris O.F. MUlier, 1 776 Leplaslerlas

= P. aporus Ludwig, 1 886 canuli Heding, 1 936
obscurus (Perrier, 1 89 1 ) clavispina Heding, 1 936
= Hexasler obscurus Perrier, 1 89 1 (Leptasterias) danica (Levinsen, 1 887)
= P. (Temnasler) hexaclis Verrill, 1 894 degerboelli Heding, 1 935
personalus Sladen, 1 89 1 floccosa (Levinsen, 1 8??)
?Plerasler sordidus Perrier, 1 894 floccosa var. crassa Heding, 1 935
= P. reduclus Koehler, 1907 (Endogenasterias) groenlandica (Steenstrup, 1 857)
pulvil/us (M. Sars, 1 86 1 ) = ASleracanthion groenlandica Steenstrup, 1 8057
Family Myxasteridae Asterias cribraria Stimpson, 1 862
Myxaster (Leplaslerias) hyperborea (Danielssen & Koren,
perrieri Koehler, 1 896 1 883)
Aslerias hyperborea Danielssen & Koren, 1 883
sol Perrier, 1 885
Pylhonasler = A sler/as normani Danielssen &Koren, 1 884
al/antidis A.H. Clark, 1948 hyperborea var. /nlermedia Heding, 1 935
(Leptaslerias ) islandlca (Levinsen, 1 887)
Order Brisingida = Aster/as mulleri var. /slandica Levinsen, 1 887
Family Brlsingidae (Leplaster/as) muelleri subsp. muelleri (M. Sars,
Brisinga 1 846)
endecacnemos Asbjemsen, 1 856 = Asteracanth/on mulleri M. Sars, 1 846

hirsula Perrier, 1 894 (Leplaster/as) muelleri subsp. ce/tica Clark &

Brisingelia Downey, 1992
coronala (0.0. Sars, 1 87 1 ) = Aster/as hispida Pennant, 1 777
= Brisinga coronala 0.0. Sars, 1 87 1 (Hexaslerias) polaris (J. Mllller & Troschel, 1 842)
= Brisingia mediterranea Perrier, 1 885 = ASleracanthion polaris J. MOller & Troschel, 1 842
Novodinia Asler/as acervata Stimpson, 1 862
homonyma Downey, 1 986 Aster/as borealis Perrier, 1 875
Brisinga e/egans Perrier, 1 885, n o n Verrill, 1 884 Marthaster/as
= Perrier, 1 894
Odinia elegans giacialis (Linnaeus, 1758)
pondina (Sladen, 1 889) Downey, 1 986 = Asteracanth/on afticanus J.MUlier, & Troschel, 1 842
Odinia pandina Siaden, 1 889 = Asler/as angulosa O.F.
MUlier, 1 788
semicoronala (Perrier, 1 885) Downey, 1 986 Aster/as echinophora Delle Chiaje, 1 827
Brisinga robusta Perrier, 1 885 =Aster/as glacialis Linnaeus, 1 75 8
= Brisinga semicoronata Perrier, 1 885 = Aster/as madelrensls Stimpson, 1 862
Asler/as rarispina Perrier, 1 875
Family Freyellidae
= /lslerias spinosa Pennant, 1777
Co/paster
M foliacea Jullien, 1 878
edwardsi (Perrier, 1 882) Downey, 1986
Sielionia webbiana d'Orbigny, 1 839
Brisinga edwardsi Perrier, 1 882
=
=

Scleraslerias
scutigeru/a Siaden, 1 889
guerne/ Perrier, 1 89 1
Freyaslera
neglecta (Perrier, 1 89 1 )
benlhophila (Sladen, 1 889) Downey, 1 986
= Aslerias edmundi Ludwig, 1 897
Freyelia benthophila Siaden, 1 889
Sto/asterias neg/ecta Perrier, 1 89 1
sexradiata (Perrier, 1 885) Downey, 1 986
richard! (Perrier, 1 882)
= Freyel/a sexradiala Perrier, 1 885
Aster/as richard/ Perrier, 1 882
luberculata (Sladen, 1 889) Downey, 1 986
Stephanasterlas
= Freyel/a tuberculala Siaden, 1 889
albula (Stimpson, 1 853)
Freyella
Asleracanlhion albulus Stimpson, 1 853
elegans (Verrill, 1 884)
= Asteracanlhion problema Steenstrup, 1855
= Brisinga elegans Verrill, 1 884
= Asterias gracilis Perrier, 1881
F. aspera Verrill, 1 895
Stichastrella
= F. bracleala Siaden, 1 889
ambigua (Farran, 1 9 1 3)
= F. laubieri Cherbonnier & Sibuet, 1973
Stichaster roseus var. ambiguus Farran, 1913
F. spinosa var. abyssicola Perrier, 1 894
rosea (O.F. MUller, 1 776)
= F. spinosa Perrier, 1 885
= Asterias rosea O.F. MOiler, 1 776
recta Koehler, 1 907
Chaelaster hermann; 1. Mllller & Troschel, 1 842
Order ForcipuJatida = Stichaster arcticus Danie1ssen & Koren, 1 8 83
Family Asleriidae Urasterias
Asteria. linckii (J. Mlllier & Troschel, 1 842)
rubens Linnaeus, 1 758 = Asteracanthion Jinckii J. MUller & Troschel, 1 842
A. clalhrala Pennant, 1 777 ASleracanthion slellionura E. Perrier, 1 869
= A. holsatica Retzius, 1 805 = Aslerias enop/a Verrill, 1 895
= /I. minula L innaeus, 1 76 1 = Aslerias gunner/ Danielssen & Koren, 1 884
= A . rubens subsp. violacea O.F. MUlier, 1 776 Family Pedicellasteridae
= A. spinosa Say, 1 825 Hydraslerias
= A. vulgar/s Packard, 1 863 sexradiala (Perrier, in Milne-Edwards, 1 882)
Cose/naslerias Pedicel/asler sexradiatus Perrier, in Milne-Edwards,
lenuispina (de Lamarck, 1 8 1 6) 1 882
= /lslerias at/anlica Verrill, 1 868 Pedicel/asler
= Asler/as savaresi Delle Chiaje, 1 827 typicus M. Sars, 1 86 1
Asler/as lenu/spina de Lamarck, 1 8 1 6 = Archasler biftons W . Thomson, 1 873
= Lylaster inaequalis Perrier, 1 894 = Astel'acanlhion palaeocrystallus Duncan & Siaden,
/casterias 1 877
panopla (Stuxberg, 1 878) Fami ly Zoroasleridae
Asterias panopla Stuxberg
Zoroaster
=

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 European Register of Marine Species

/u/gens Thomson, 1 873 Order Ophiurida

Family Labldiasteridae Family Opbiotricbidae
Coronaster Ophiolhrix
volsel/alus (Sladen, 1 889) fragilis (Abildgaard, in O.F. MUller, J 789)
=Asterias (Slo/aslerias) valsella/us Sladen, 1 889 = Asteria cuvieri Delle Chiaje, 1 828
=C. antonii Perrier, 1 894 Aslerias echinala Delle Chiaje, 1 822-29
Family Neomorphasteridae = Asteriasfragifis Abildgaard, in O.F. MUlier, 1 789
Neomorphasler Aslerias penlaphylla Pennant, 1 777
margaritaceus (Perrier, in Milne-Edwards, 1 882) = Aslerias rllbra Delle Chiaje, 1 8??
A.M. Clark & Downey, 1 992 = Ophiocoma minula Forbes, 1 839
Calycaster monoecus Perrier, 1 89 1 = Ophiocoma rosula Forbes, 1 839
=N. eustichus Siaden, 1 889 = O. alopecurus I. MUller & Troschel, 1 842
=N. par/aiti Koehler, 1 896 = o. /erllssacii J. MUller & Troschel, 1 842
= Pedicellaster margarilaceus Perrier, in Mi lne-Edwards, = O. lusitanica Ljungman, J 872
1 882 = 0. rammefsbergii J. MUlier & Troschel, 1 842
=Stichasler talisman; Perrier, 1 88S 0. rubra Ljungman, J 872
= Ophiura qllinquemaculala Delle Chiaje, 1 828
Class Ophiuroidea indigna Koehler, 1 906
Order Phrynophiurida luetken; Wyville Thomson, 1 873
macu/ala Ljungman, 1 872
Family Ophiomyxidae
= 0. indllcla Koehler, 1 906
Aslrogeron
supinus (Lyman, 1 8 83) Family Opbiactidae
Ophiogeron supinus Lyman, 1 883 Hislampica
Ophiogeron dllplicata (Lyman, 1 875)
edenlulus Lyman, 1 878 Ophiaclis
Ophfomyxa abyssicola (M. Sacs, 1 86 1 ) Ljungman, 1 866
penlagona (de Lamarck, 1 8 1 6) = ?Ophiactis cuspidala Lyman, 1 882
= Ophfura penlagona de Lamarck, 1 8 1 6 = Amphiura abyssicola M. Sars, 1 86 1
serpentaria Lyman, 1 883 = O. corallicola Koehler, 1 896
Ophiophrixus 0. echinala Koehler, 1 898
spinosus (Storm, 1 8 8 1 ) = O. poa Lyman, 1 882
= Ophiobyrsa hyslricis Lyman, 1 8 83 balli (W. Thompson, 1 840)
Ophfobyrsel/a hyslricis (Lyman, 1 883) = Ophiocoma balli W Thompson, 1 840
Ophioscolex = Ophiocoma goadsiri Forbes, 1 840 (Nov.)
g/acia/is J. MUller & Troschel, 1 842 canotia Lyman, 1 882
purpureus DUben & Koren, 1 846 /ymani Ljungman, 1 872
?Ophiosco/ex quadrispinus Verrill, 1 88 5 nidarasiensis Mortensen, 1 920
profundi LU!ken & Mortensen, 1 899
Order Euryalina
savignyi (1. MUller & Troschel, 1 842)
Family Asteronycbidae O. brocki de Loriol, 1 893
Asleronyx = O. krebsii LUtken, 1 857
loveni J. MUlier & Troschel, 1 842 Ophiolepis savignyi J . MUlier & Troschel, 1 842
= A. cooped Bell, 1 909 virens (M. Sars, 1 8S7)
= A. dispar LU!ken & Mortensen, 1 899 = Amphiura virens M. Sars, 1 857
A. /ocardi Koehler, 1 896 Ophiopholis
= Ophiuropsis Iymani Studer, J 884 aculeala (Linnaeus, 1 767)
Astrodia = Ophiocama bellis Forbes, 1 839
lenuispina (Verrill, 1 884) Ophiolepis sc% pendrica 1. MUlier & Troschel, 1 842
= A. bispinasa Koehler, 1 922 Polypholis echinala Duncan, 1 8??
Family Gorgonocepbalidae
Ophiopus
Astrosparlus arcticus Ljungman, 1 867
medilerraneus (Risso, 1 826) Ophiaregma abyssorum G.O. Sars, 1 872
= Euryale arborescens L. Agassiz, 1 839 Family Amphiuridae
= Euryale medilerraneus Risso, 1 826 Amphichilus
Gorgonocepha/us verrucosus Grube, 1 840 da/eus (Lyman, 1 882) Matsumoto, 1 9 1 7
Gorgonocephalus Amphiura dalea Lyman, 1 882
arclicus Leach, 1 8 1 9 Amphioplus
= Astrophylon agassizi Stimpson, 1 853 verrilli (Lyman, 1 879)
capu/medusae (Linnaeus, 1 758) Silox pu/vinus Cherbonnicr & Sibuet. 1 972
= Aslerias caput-medusae Linnaeus, 1 758 Amphipholis
Astrophyton linckii I. MUller & Troschel, 1 842 instructa Koehler, 1 906
= Astrophyton sculalum Fleming, 1 828 murmanica D'yakonov, 1 929
eucnemis (I. MUlier & Troschel, 1 842) Lyman, 1 882 squamata (Delle Chiaje, 1 829)
= Astrophyton eucnemis I. Milller & Troschel, 1 842 A. lineala Ljungman, 1 872
= Astrophylon malmgren; Danielssen & Koren, 1 877 A. tenera LUtken, 1 8??
= G. caryi Lyman, 1 860 Amphiura e/egans (Leach, 1 8 1 5)
G. japonicus Dliderlein, 1907 = Asterias squamala Delle Chiaje, 1 829
/amarckii (I. MUller & Troschel, 1 842) Lyman, 1 882 =Axiognalhus squamala (Delle Chiaje)
= Astrophyton lamarcki J. MUlier & Troschel, 1 842 torelli Ljungman, 1 872
Family Asteroschematidae
Amphiura
Asteroschema abyssorum Norman, in Jeffreys, 1 876
inornatum Koehler, 1 9?? =?A. digna Koehler, 1 906
Ophiocreas bellis Lyman, 1 879
oedipus Lyman, 1 879 bellis var. Irilonis Hoyle, 1 884

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 Emblow and White 200 I

borealis (G.O. Sars, 1 87 1 ) Ljungman, 1 872 nolata Koehler, 1 906

= Ophiope/tis borealis G.O. Sars, 1 8 7 l opercu/aris (Koehler, 1 909)
(Acrocnida) brachiata (Montagu, 1 804) = Ophiolimna opercularis Koehler, 1 909
= Amphiocnida brachiata (Montagu) parcita Koehler, 1906
= A. neapolitana M. Sars, 1 857 setosa (Retzius, I S05)
= Asterias brachiala Montagu, 1 804 Ophiura selosa Retzius, 1 805
cherbonnieri Guille, 1 972 smitti Ljungman, 1 872
chiajei Forbes, 1 843 slmulans Koehler, 1 896
defamare; Cherbonnier, 1 958? O. composita Koehler, 1 906, non Koehler, 1 904
denticulata Koehler, 1 89 = 0. composila Mortensen, 1 927
filiformis (O.F. MUlier. 1 776) spec/abi/is G.O. Sars, I S 7 1
= Asleriasfiliformis O.F. Mill ler, 1 776 spinosella Mortensen, 1 9 3 3
fragilis Verrill, 1 885 valenciennesi Lyman, 1 879
= A. denticulala Koehler, 1 898 Ophiambix
grandisquama Lyman, 1 869 devaneyi Paterson, 1 985
= A. iris Lyman, 1 879 meteoris Bartsch, 1 983
= A. josephinae Ljungman, 1 872 Ophiochondrus
= A. longispina Koehler armalus (Koehler, 1 909)
griegi Mortensen, 1 920 = Ophioplus armalus Koehler, 1 909
incana Lyman, 1 879 Ophiocomina
mediterranea Lyman, 1 882 nigra (Abildgaard, in O.F. Milller, 1 789)
olleri Ljungman, 1 872 Asterias nigra Abildgaard, in O.F. Milller, 1 789
= A. grandis Koehler, 1 896 Ophiacanlha danae Mortensen, 1 933
palmeri Lyman, 1 882 = Ophiocoma granuiata Forbes, 1 839
richardi Koehler, 1 906 Ophiocoma nigrll (Abildgaard)
sarsi Ljungman, 1 872 = Ophiocoma nilssoni J. MUlier & Troschel, 1 842
(Ophiope/tis) securigera (Dilben & Koren, 1 846) = Ophiocoma raschi G.O. Sars, IS??
Ophiopeltis securigera DUben & Koren, 1 846 Ophiolebes
(Acrocnida) sem/squamata (Koebler) bacata Koehler, 1909
senega/ensis Madsen, 1 970 retecta (Koehler, 1 896)
stepanovi D'yakonov, 1 954 Ophioscolex retectus Koehler, 1 896
= A. stepanovi Chernyavskii, 1 86 1 (nomen nudum) Ophiomitra
A. stepanovi Lyman, 1 869 (nomen nudum) hamula Mortensen, 1 933
sundevalli O. Millier & Troschel, 1 842) Ophiopsila
= Ophiofepis sundevalli J. Milller & Troschel, I S42 annulosa (M. Sars, 1 859)
Monamphiura Ophianoplus annulosus M. Sars, 1 859
apicula (Cherbonnier, 1 957) aranea Forbes, 1 845
= Amphiura apicula Cherbonnier, 1 957 = Ophianoplus marmoreus M. Sars, 1 8??
Paramphlura alT. guineensls (Koehler, 1 9 1 4)
punelala (Forbes, 1 84 I ) Ophiocamax
Ophiaeoma punetata Forbes, 1 84 1 dominans Koehler, 1 9??
Ophiomifrella
Family Ampbilepididae
clavigem (Ljungman, 1 865)
A mphilepis
?Ophiomitrella cordifera Koehler, 1909
norveglca (Ljungman, IS65)
Ophiuclis clavigera Ljungman, I S65
= Amphiura norvegica Ljungman, 1 865
=

Ophio/ebes acanellae Verrill, I S85

ingolfiana Mortensen, 1 93 3
=

Ophiolebes claviger (Ljungman, 1 865)

Order Laemophiurina globulifera (Koehler, 1 896)
Family Ophiacanlbidae Ophiop/inlhuca
Ophlacantha carduus (Lyman, 1 882)
abyssicola G.O. Sars, 1 87 1 chelys (Lyman, 1 882)
O. mil/espina Verri ll, 1 879 Ophiothamnus
angolensis Koehler, 1 923 ajJinis Ljungman, 1 872
anomala G.O. Sars, 1 87 1 = Ophioleda minima Koehler, 1 906
?Ophiacantha nodosa Lyman, I S?? Ophioplinthaca occlusa Koehler, 1 907
arislata Koehler = O. minima Matsumoto, 1 9 1 7
bidentata (Retzius, 1 805) charUs H.L. Clark, 1 9 1 5
Asterias bidentala Retzius, 1 805 O. gracilis (Verrill, 1 885), non (Studer, 1 822)
= 0. fratema Verrill, 1 885 Ophlolimna
0. groenlandica J. MUller & Troschel, 1 844 bairdi (Lyman, 1 883) Verril l , 1 899
= O. hibernica Farran, 1 9 1 3 = Ophiacantha bairdi Lyman, 1 883
= 0. spinulosa J . MUlier & Troschel, 1 842 Ophiomedea
brevispina Koehler, 1 898? duplicata Koehler, 1906
composifa Koehler, 1 907 Ophioprium
cra.fsidens Verrill rosea (Lyman, 1 879) Baker & Devaney, 1 9 8 1
cuspidata Lyman, 1 882 Ophiacanlha rosea Lyman, 1 879
(Ophiotreta) danae Mortensen, 1 933 = Ophiacantha t/'lmcata Koehler, 1 930
0. valenciennesi Lieberkind, 1 929, non Lyman Ophiotoma
decipiens Koehler, 1 906 bartlelli (Lyman, 1 8??)
densa Farran, 1 9 1 3? coriacea Lyman, 1 883
enopia subsp. velerna (Koehler, 1 907) Ophiolrema
hirsllta Lyman, 1 875 alberti Koehler, 1 896
lineata Koehler, 1 909 Ophiomyces
mesembria H.L. Clark, 1 9 1 5 frutectosus Lyman, 1 869
nodosa Lyman, 1 882 grandis Lyman, 1 878

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 European Register of Marine Species

Order Chilophiurina = Ophiozona sincera Koehler, 1 906

tjalftana (Mortensen, 1 9 1 3) Matsumoto, 1 9 1 5
Family Ophiuridae
Ophiozona tjalftana Mortensen, 1 9 1 3
Amphiophiura
Ophiura
bullata (Wyville Thomson, 1 873) Matsumoto, 1 9 1 5
albida Forbes, 1 839
= Ophioglypha abdita Koehler, 1 909
africana (Koehler)
Ophioglypha bul/ala Wyville Thomson, 1 873
= Ophiocten africana Koehler
bul/ala subsp. eonvexa (Lyman, J 878)
(Dictenaphiura) carnea LUtken, 1 858 ex M . Sars MS
= Ophioglypha convexa Lyman, 1 878
clemens (Koehler, 1 906)
bullara subsp. viljazi Litvinova, J 971
= Ophioglypha clemens Koehler, 1 906
= A. viljazi Litvinova, 1971
jlagellala (Lyman, 1 878)
saurura (Verrill, J 894)
Gymnophiura caerulescens Lmken & Mortensen' 1 899
= Ophioglypha aspera Koehler, 1 898
=

= Ophiog/yphaj/agellata Lyman, 1 878

= Ophioglypha saurura Verrill, J 894
grubei Heller, 1 863
Anlhophiura
imprudens (Koehler, 1 906)
ingolft Fasmer, 1 930
Ophioglypha imprlldens Koehler, 1 906
Aspidophiura
inermis (Lyman, 1 882)
minilia (Lyman, 1 878)
= Ophiog/ypha inermis Lyman, 1 882
= Ophiog/ypha minuta Lyman, 1 878
(Ophiuroglypha) irrorata (Lyman, 1 878)
Homophiura
= Ophioglypha grandis Verrill, 1 894
abyssorurn (Lyman, 1 883)
= Ophioglypha involuta Koehler, 1 899
abyssorum var. africana (Madsen, 1 947)
Ophioglypha irrorala Lyman, 1 87 8
Homalophiura abyssorum var. afrieana Madsen, 1 947
= Ophioglypha orbiculala Lyman, 1 882
lessellata (Verrill, 1 894)
Ophioglypha lumu/osa Liitken & Mortensen, 1 899
= Homalophiura tessel/ata (Verrill, 1 894)
=

(Ophiuroglypha) irrorata subsp. concreta (Koehler,

Ophioglypha tessellala Verrill, 1 894
1 90 1 )
Ophiocten
= Ophiog/ypha concreta Koehler, 1 90 1
abyssicolum von Marenzeller, 1 8??
(Ophiuroglypha) irrorata subsp. po/yacantha
affinis (LUtken, 1 858) Sumida, Tyler, Gage &
Mortensen, 1 93 3
N0rrevang, 1 998
Homalophiura mullispina Cherbonnier & Sibuet, 1 972
Ophiura affinis Liitken, 1 858
(jungmani (Lyman, 1 878)
cenlobi Paterson, Tyler & Gage, 1 982
= Ophioglypha /epida Lyman, 1 882
gracilis (G.O. Sars, 1 87 1 )
Ophiog/ypha (jungmani Lyman, 1 87 8
0. serieeum auctt., non (Forbes, 1 852) (p.p.)
= Ophiog/ypha Ihouleli Koehler, 1 896
= Ophiog/ypha gracilis G.O. Sars, 1 8 7 1
madseni (Belyaev, 1 972)
= Ophiog/ypha signala Verrill, 1 882
= Homalophiura madsen; (Belyaev, 1 972)
Ophiura signata (Verrill, 1 882)
(Ophiurog/ypha) mundata Koehler, 1 906
haslatum Lyman, 1 878
nilida Mortensen, 1 93 3
o.phioclen longispinum Koehler, 1 898
ophiura (Linnaeus, 1 758)
= O. /atens Koehler, 1 906
= Asterias ophiura Linnaeus, 1 75 8
= O. pacifieum Latken & Mortensen, 1 899
0. cWaris (Linnaeus, 1 766)
sculalum Koehler, 1 898
= 0. ciliala (Retzius, 1 783) non (O.F. MUlier, 1 776)
serieeum (Forbes, 1 852)
= O. lacerlosa (Pennant, 1 777)
= 0. kroeyeri Liltken, 1 85 5
O. lexlurala de Lamarck, 1 8 1 6
Ophiura sericea Forbes, 1 852
robllsla (Ayres, 1 8 5 1 )
Ophioclenella
= Ophiolepis robus/a Ayres, 1 8 5 1
ades Tyler, Paterson, Sibuet, Guille, Murton &
O. /ascicu/ala Forbes, 1 854
Segonzac, 1995
= O. squamosa LUtken, 1 85 9
Ophio/epis
sarsii LUtken, 1 858
paucispina ?
violaine (Cherbonnier & Sibuet, 1 972)
Ophiomisidium
= Homalophiura vio/aine Cherbonnier & Sibuet ' 1 972
pulchellum (Wyville Thomson, 1 8??)
Ophiurolepis
Ophiomusium pulchellum Wyville Thomson, 1 8??
inornata (Lyman, 1 878) Paterson, 1 985
Ophiomusium
= Homalophiura inornala (Lyman, 1 878)
africanum Koehler, 1 909
Ophioglypha indivisa LUtken & Mortensen, 1 899
/ymani Wyville Thomson, 1 873
= Ophioglypha inornata Lyman, 1 878
Ophiophycis
Slegophiura
mirabilis Koehler, 1 90 I
nodosa (LUtken, 1 858) Matsumoto, 1 9 1 7
Ophiopleura
= Ophiura nodosa Liltken, 1 85 8
aurantiaca (Verrill, 1 882) Verrill, 1 882
stuwilzii (LUtken, 1 858) Matsumoto, 1 9 1 7
Ophioglypha aurantiaea Verrill, 1 882
= Ophiura slllwilzii LUtken, 1 85 8
borealis Danielssen & Koren, 1 877
Uriopha
= Luetkenia arc/iea Duncan, 1 878
ios Paterson, 1 980
Ophiosphalma
armigerum (Lyman, 1 878) H.L. Clark, 1 9 4 1 Family Ophioleucidae
= Ophiomusium armigerum Lyman, 1 878 Ophiernus
planum (Lyman, 1 878) H.L. Clark, 1 94 1 adspersus Lyman, 1 883
= Ophiomusium planum Lyman, 1 878 vallincola Lyman, 1 878
Ophiotjalfa O. abyssalis Koehler, 1 896
vivipara Mortensen, 1 9 1 3 Ophiocrates
Ophiotypa interval/us Madsen, 1 947
simplex Koehler, 1 897 secundus Koehler, 1 906
Ophiozonella Ophiopyren
molesfa (Koehler, 1 906) striatum Mortensen, 1 933
Ophiozona molesfa Koehler, 1 906 Family Ophionereididae
sineera (Koehler, 1 906) Ophiochiton

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Costello, Emblow and White (eds), 200 1

solutus Koehler, 1 906 grima/dii Koehler, 1 897

lern;spinus Lyman, 1 883 Tromikosoma
= 0. grandis Verrill, 1 884 koehleri Mortensen, 1 903
O. tenuispinus Hoyle, 1 884
Order Diadematoida
Ophionereis
reticulata (Say, 1 825) Family Aspidodiadematidae
= Ophiura reticulata Say, 1 825 Aspidodiadema
jacobyi A. Agassiz, 1 883
Family Ophiodermatidae
Plesiodiadema
Bathypectinura
anlil/arum (A. Agassiz, 1 883)
heros (Lyman, 1 879)
Aspidodiadema antil/arum A. Agassiz, 1 883
=
= B. elata (Koehler, 1 906)
= Pectinura elala Koehler, 1 906 Family Diadematidae
= Pectinura heros Lyman, 1 879 Cenlrostephanus
Cryptopelta longispinus (Philippi, 1 845)
brcvispina (Ludwig, 1 879) Cidaris (Diadema) longispina Philippi, 1 845
Ophioconis brevispina Ludwig, 1 879 Diadema
Ophiarachnella anlil/arum Philippi, 1 845
scmicincla (Studer, 1 8??) Cidaris {Diadema} antil/arum Philippi, 1 845
= Peetinura semicincta Studer, 1 8?? Order Pedinoida
Ophioconis
Family Pedinidae
forbesi (Heller, 1 863)
Caenopedina
Pectinuraforbesi Heller, 1 863
cubensis A. Agassiz, 1 869
vivipara Mortensen, 1 9??
Ophioderma Order Salenoida
longicauda (Retzius, 1 805) Family Saleniidae
Asterias longicauda Retzius, 1 805 Sa/enia
= O. lacertosum Lamarek, 1801 patterson; A. Agassiz
Ophiura laevis Lyman, 1 865 Salenocidaris
Pectinura profundi subsp. profundi (Duncan, 1 877)
vestita Forbes, 1 843 = ?Sa/enia hastigera A. Agassiz
Class Echinoidea Salenia hastigera subsp. aeuminala Mortensen, 1935
Sa/enia profundi subsp. oec/usa Mortensen, 1934
=

Order Cidaroida
= Sa/enia profundi Duncan, 1 877
Family Cidaridac profundi subsp. brevispina Markov, 1988
Cidaris varispina A. Agassiz, 1 869
cidaris (Linnaeus, 1 758) Order Arbacioida
= C. borealis DUben & Koren, 1 846
= C. papil/ala Leske, 1778 Family Arbaciidae
= Cidarites hyslrix de Lamarck, 1 8 1 6 Arbaeia
= Dorocidaris abyssicola A . Agassiz, 1 8697 lau/a (Linnaeus, 1 758)
= Echinus cidaris Linnaeus, 1 758 Cidaris puslu/osa Leske, 1 778
Iribu/oides de Lamarck, 1 8 1 6 Echinus aequitubercu/atus de Blainville, 1 825
=

Poriocidaris Echinus /ixu/a Linnaeus, 1 758

purpurala (Wyville Thomson, 1 875) Echinus neapolitanus Delle Chiaje, 1 82 5
=

= Porocidaris gracilis Siaden, 1891 Arbaciel/a

= Porocidaris purpurata Wyville Thomson, 1875 e/egans Mortensen, 1 9 1 0
Siereocidaris Coe/op/eurus
ingolfiana Mortensen, 1 903 floridanus A. Agassiz, 1 872
Stylocidaris Habrocidaris
affinis (Philippi, 1 845) scutata (A. Agassiz, 1 880)
= Cidaris affinis Philippi, 1 845 Podoeidaris
Cidaris stokesii L. Agassiz & DesoT, 1 846-47 scu/pla A. Agassiz

Order Ecbinotburoida Order Temnopleuroida

Family Echinotburidae Family Temnopleuridae

Calveriolloma Genocidaris
feneslralum (Wyville Thomson, 1872) macula/a A. Agassiz, 1 869
= Araeosoma violaceum Mortensen, 1 903 = Arbacina pal/aryi Gauthier, 1 897
= Calveriafeneslrala Wyville Thomson, 1 872 Temnechinus macula/us A. Agassiz, 1 869
hystrix (Wyville Thomson, 1 872) Hypsiechinus
= Aslhenosoma reynoldsii A. Agassiz, 1880 coronaWs Mortensen, 1903
Calveria hyslrix Wyvil le Thomson, 1 872 Trigonocidaris
Hygrosoma a/bida A. Agassiz
pelersii (A. Agassiz, 1 880) Family Toxopneusfidae
= Phormosoma luculenlum Koehler, 1 896, non A. Sphaerechinus
Agassiz, 1 8?? granularis (de Lamarck, 1 8 1 6)
= Phormosoma peters;; A. Agassiz, 1 880 Echinus (Toxopneusles) brevispinosus Risso, 1 826
Phormosoma uranus Koehler, 1 898, non Wyville = Echinlls granularis de Lamarck, 1 8 1 6
Thomson, 1 877 S. rosells Russo, 1 893
uranus (Wyville Thomson, 1 877)
Order Ecbinoida
= Phormosoma uranus Wyville Thomson, 1 877
Phormosoma Family Echinidae
placenta Wyville Thomson, 1 874 Echinus
P. sigsbei A. Agassiz, 1 883 aculus de Lamarck, 1 8 1 6
Sperosoma ?Echinus depressus G.O. Sars, 1 871

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 European Register of Marine Species

?Echt'nus micros/oma Wyville Thomson, 1 875 alT. alcocki Koehler, 1 9 1 4

?Echinus norvegiclIs Duben & Koren, 1 846 =P. miranda auctt. non A . Agassiz, 1 869
= ?Echinus rarispinus G.O. Sars, 1 872 jeffreysi Wyville Thomson. 1 873
E. flemingU Forbes, 1 84 1 wandeli Mortensen, 1 905
E. sardicus Delle Chiaje, 1 84 1
Family Urechinidae
ajfinis Mortensen, 1 903 Urechinus
alexandri Danielssen & Koren, 1 883 naresianus A. Agassiz, 1 88 1
elegans Duben & Koren, 1 846
Family Plexechinidae
?Echinus wallisi A. Agassiz, 1 883
Plexechinus
esculentus Linnaeus. 1 75 8
hirsutus Mortensen, 1 905
E. aurantiacus de Blainville. 1 82 5

E. globiformis d e Lamarck. 1 80 I Family Calymnidae

= E. pseudomelo de Blainville. 1 825 Calymne
E. quinqueangulatus de Blainville, 1 825 relicta Wyville Thomson, 1 877?
E. quinqueangulosus de Blainville, 1 834 Order Spatangoida
E. schwartz;; Nilsson & Holst. 1 8 1 7

E. sphaera O.F. MUlier, 1 776 Family Hemiasteridae

= E. subangulosus de Blainville, 1 826 Hemiaster
= E. violaceus de Blainville, 1 825
expergitus Loven, 1 874
= H. gibbosus A. Agassiz. 1 88 1
esculentus var. glacialis D'yakonov, 1 923
melD Lamarck, 1 8 1 6 H. mentzii A . Agassiz. 1 883
= H. zonatus A. Agssiz, 1 881
tenuispinus Norman. 1 868
E. esculen/us var. tenuispinus Norman, 1 868 Family Spatangidae
Paracenlrotus Spatangus
lividus (de Lamarck, 1 8 1 6) inermis Mortensen, 1 9 1 3
= Echinus lividus de Lamarck, 1 8 I 6 purpureus (O.F. MUlier, 1 776)
= Echinus purpureus Risso, 1 826 = Spatagus purpureus O.F. MUlier, 1 776

Echinus saxatilis Tiedemann, 1 8 1 6 S. 'meridionalis Risso, J 825

= Echinus vulgaris de Blainville, 1 825 = S. reginae Gray, 1 85 5
Psammechinus = S, spinosissimus Desor, 1 847
microtuberculatus (de Blainville, 1 825) Heller. 1 86 8 rasch/ Loven, 1 869
Echinus microtuberculatus d e Blainville, 1 825 = S, meridionalis Norman, 1 8?? non Risso, 1 825
miliaris (P.L. S , MUlier, 177 1 )
Family Schizasteridae
Echinus miliaris P.L.S. MUlier, 1 766
Brisaster
= Parechinus miliaris Mortensen, 1 903
fragilis (Dilben & Koren, 1 846)
Family Strongylocentrotidae Brissusfragilis Duben & Koren, 1 846
Strongylocentrotus Schizaster fragilis (DUben & Koren)
=

droebachiensis (O.F. MUller, 1 776) Schizaster

= Echinus drobachiensis O.F. MUlier. 1 776 canaliferus (de Lamarck. 1 8 1 6)
= Echinus granularis Say, 1 82 6 = Spatangus canaliferus de Lamarck, 1 8 1 6
= Echinus neglec/us d e Lamarck, 1 8 1 6
Family Brissidae
S. chlorocentrotus Brandt, 1 835
Brissopsis
= Toxopneustes pictus Norman, 1 8??
atlantica Mortensen, 1 907
pallidus (G.O. Sars, 1 87 1 )
atlantica var. mediterranea Mortensen, 1 9 1 3
= ?Strongylocentrotus granular;s (Say, 1 827)
Iyrifera (Forbes, 1 84 1 )
= S. echinoides A. Agassiz & H,L. Clark, 1 907
= B. parma Val., 1 8??

S. pic/us (Norman, 1 868) Brissus Iyrifer Forbes, 1 84 1

= S, sachalinicus Clark, 1 9 1 2
= Brissus pulvinatus Philippi, 1 845

Order Clypeasteroida = Schizaster incer/us Aradas, 1 850

Brissus
Family Fibuillriidile
unicolor (Leske, 1 778)
Echinocyamus
= B, brissus auctt.
grandiporus Mortensen. 1907
B. scil/ae L. Agassiz, 1 836
macros/omus Mortensen, 1907
= Spatangus brissus subsp. unieolor Leske, 1 77 8
pusillus (O.P, MOiler, 1 776)
Spatangus columbaris d e Lamarck. 1 8 1 6
= E, angulosus Leske. 1 778

Plagiabrissus
= E. parthenopaeus Costa, 1 869
ajfinis ?
E. speciosils Costa, 1 869
costai (Gasco. 1 876)
= Echinus minulus Omelin, 1 790
= Metalia costae Gasco, 1 876
= Fibularia equina Aradas, 1 850
= Rhabdobrissus perrieri Bernard
Fihularia tarentina de Lamarck, 1 8 1 6
= Spa/agus pusillus O.F. MUlier, 1 776 Family Loveniidae
Echinocardium
Order Cassiduloida
cordatum (Pennant, 1 777)
Family Neolampadidae = Amphidetus kiirtzii Girard. 1 852

Neolampas = E. sebae Gray. 1 8??

rostellata A. Agassiz, 1 869 Echinus cordatus Pennant, 1 777

= S pa/angus arcuarius de Lamarck, 1 8 1 6
Order Pourtalesioida
fenauxi Pequignat, 1 963
Family Pourtalesiidae flavescens (O.P. MUlier. 1 776)
Echinosigra = A mphidotus roseus Forbes, 1 84 1

phiale (Wyville Thomson. 1 873) Mortensen. 1 907 = Spalagus flovescens O.F. MUlier. 1 776

= E, paradoxa (Mortensen, 1 905) Mortensen. 1907

Spatangus ovatus Leske, 1 778
= Pourtalesia paradoxa Mortensen, 1 905
mediterranellm (Forbes. 1 844)
= Pourtalesia phiale Wyville Thomson, 1 873
Amphidetus mediterraneus Forbes, 1 844
Pourtalesia

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Costello, Emblow and White (eds), 200 1

= E. gibbosus L. Agassiz & Desor, 1 847 Cucumaria communis Forbes, 1 84 1

marlenseni Thiery, 1 909 = Cucumaria nobilis Ludwig, 1 874
= ?Echinocardium laevigasler A. Agassiz, 1 8?? Cucumaria perspicua Ludwig, 1 874
?Echinacardium pennatifidum Koehler, 1 898, nOli = Holo/huria drummondii Thompson, 1 840
Nonnan, 1 868 Thyone porllocki Forbes. 1 84 1
= E. intermedium Mortensen, 1 907 = T. diibeni Nonnan, 1 869
pennatifidum Nonnan, 1 868 Family Cucumariidae
= E. gibbosus Barrett, 1 857 non L. Agassiz & Desor, Abyssocucumis
1 847 abyssorum (Theel, 1 886)
Family Aeropsidae = Cucumaria abyssorum auctt., non Theel. 1 886
Acesle Cucumaria abyssorum Theel. 1 886
bellidifera Wyville Thomson, 1 877 = Cucumaria sluiter; Ohshima, 1 9 1 5
Aeropsis = Staurocucumis ingolfl Clark & Deichmann, 1 936
rastrata (Wyville Thomson, 1 877) Mortensen, 1 907 Aslia
= Aerope rostra/a Wyville Thomson, 1 877 le/evrei (Barrois, 1 882)
Family Asterostomatidae = Cucumaria leJevrei Barrois, 1 882
Palaeotropus
Cucumaria normani Allen & Pace, 1 904
josephinae Loven, 1 8??
= Hololhl/ria monlagui Fleming, 1 828 (p.p.)
= Palaeobrissus hilgardi A. Agassiz Cucumaria
arcuala (Herouard, 1 92 1 )
= P. hirondellei Koehler
Peripatagus
= Psolidium arcua/um Herouard, 1 92 1
cinctus Koehler
compressa (R. Perrier, 1 898)
Ocnl/Ii compressus R. Perrier, 1 898
Class Holothuroidea /rondosa (Gunnerus, 1 767)

Order Dendrochirotida C. assimilis DOben & Koren, 1 846

= Holo/huria /rondosa Gunnerus, 1 767
Family Psolidae Hololhuriafucicola Forbes & Goodsir, 1 839
Psolidium =Holothuria grandis Forbes & Goodsir, 1 839
complana/um Cherbonnier, 1 969 Hololhuria pen/acles Linnaeus, 1 767
Psolus paraglacialis Heding, 1 942
fabricii (DUben & Koren, 1 846) Havelockia
Cuvieria/abricii DUben & Koren, 1 846 inermis (Heller, 1 868)
hypsinotus Heding, 1 942 Thyone elegans Nonnan, 1 869
nummularis R. Perrier, 1 899 = Thyone inermis Heller,1 868
opercula/us (Pourtales, 1 868) Urox;a auranliaea Costa, 1 869
Cuvieria operculala Pourtales, 1 868 Neocnus
phan/apus (Strussenfelt, 1 765) Jaeger, 1 833 incubans Cherbonnier, 1 972
= Hololhuria phan/apus Strussenfelt, 1 765 Ocnus
= P . granulalus Ayres, 1 85 1 glacialis (Ljungman, 1 879)
P. laeviga/us Ayres, 1 85 1 = Cueumaria glacialis Ljungman, 1 879
= P. regalis Verrill, 1 866 grubei (von Marenzeller, 1 874)
pour/ales;; Theel, 1 886 = Cucumaria dicquemarii M. Sacs. 1 857
squama/us (O.F. MUller, 1 776) L[Jtken, 1 857 Cucumaria grubei von Marenzeller. 1 874
Holo/huria squamala O.F. MOller, 1 776 = Pseudocnus grubei Panning, 1 949
= P. valvallls Ostergren, 1 904 koellikeri (Semper, 1 868)
lesselalus Koehler, 1 896 =Cucumaria Mllikeri Semper, 1 868
Family Phyllophoridae Pseudocnus dubiosus (Semper) Panning, 1 962
Ekmania =Pseudocl1us kollikeri Panning. 1 949
bar/hii (Troschel, 1 846) lacleus (Forbes & Goodsir, 1 839)
Orcula barlhii Troschel, 1 846 =Cucumaria mos/erensis Grieg, 1 889
= Orcula luminosa Lampert, 1 885 Hololhuria laclea Forbes & Goodsir. 1 839
Thyonidium pellucidum DUben & Koren, J 846 pe/iti (Cherbonnier, 1 958)
Euthyonidiella Ludwigia petiti Cherbonnier. 1 958
allantica (Ludwig & Heding, 1 935) planei (Brandt, 1 835)
= Pseudocucumis a/lanlica Ludwig & Heding, 1 935 ?Holothuria brunneus (W. Thompson, 1 840. ex Forbes
Neocucumis MS), non Chamisso & Eysenhardt, 1 82 1
arlantica (Ludwig & Heding, t 935) ?Ocnus brunneus Forbes & Goodsir, in Forbes, 1 84 1
marioni (von Marenzeller, 1 878) =Cladodactyfa plonci Brandt, 1 835
= Cucl/maria marioni von Marenzeller, 1 878 syracusanus (Grube, 1 840) Panning, 1 949
Pseudocucumis marioni Ludwig. 1 874 =Cladodactyla syracusana Grube, 1 840
Neopenladactyla Cucumaria hyndmanni W. Thompson, 1 840
mix/a (Ostergren, 1 898) Deichmann, 1 944 =Cucumaria syracusana auctl.
= Pseudocucllmis cueno/i Koehler & Vancy, 1 905 Pseudocnus syracusanus auctt.
Pseudocucumis mix/a Ostergren, 1 898 Paracucumaria
Phyllophorus mauri/anica (Herouard, 1 929)
drachi Cherbonnier & Guille, 1 968 = Cucumaria mauritanica Herouard, 1 929
granulatus (Grube, 1 840) thalassae Cherbonnier, 1 969
Psolus granulalus Grube, 1 840 Pawsonia
(Phyllophorus) pedinaequalis Cherbonnier, 1 969 saxicola (Brady & Robertson, 1 87 1 )
(Phyllophorus) IIrna Grube, 1 840 = Colochirus l!lcazei Herouard
= Holothuria penicillus Delle Chiaje, 1 828 Cucumaria saxicola Brady & Robertson, 1871
Thyonidium = Hololhuria mOlllagui Fleming,1 828 (p.p.)
hyalinum (Forbes, 1 84 1 ) aff. saxicola Garrido (email check-list ofCanarian
= Holo/huria pellucida Fleming, 1 828, non Vahl, 1 806 echinoderms)
drummondii (Thompson, 1 840) Penlamera

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 European Register of Marine Species

calcigera (Stimpson, 1 85 1) Slichopus moseleyi Theel, 1 886

=

= Cueumaria koreni LOtken, 1 857 pourlalesii (Theel, 1 886)

Penlacta calcigera Stimpson; J 85 1 B. bipartitus Herouard, 1 923
=

Pseudolhyone Stichopus pourtalesii Theel, 1 886

=

furnesli Cherbonnier, 1 969 lizardi (Theel, 1 882)

raphanus (Dilben & Koren, 1 846) B.fallax Ostergren, 1 896
=

Thyone poucheri Barrois, 1 882 = B. replans R. Perrier, 1 902

= Thyone raphanus DUben & Koren, 1 846 = Stichopus lizardi Theel, 1 882
seulponea Cherbonnier, 1 958 Benlhothuria
serrifera (Ostergren, 1 898) Jensenius Madsen & funebris R. Perrier, 1 898
Hansen, 1 994 ?Bentho/huria valdiviae Heding, 1 940
=

= Thyone serrifera O stergren, 1 898 Gephyrothuria

Slereoderma alcocki Koehler & Vaney, 1905
incerta Cherbonnier, 1 969 = G. europeensis Herouard, 1 923
kirchsbergi (Heller, 1 868) Mesothuria
= Cucumaria kirchsbergi Heller, 1 868 bifurcata Herouard, 1 906
Gcnus? kirchsbergi von Marenzeller, 1 874 candelabra Herouard, 1 923
Thyone cathedralis Heding, 1 940
cherbonnieri Reys, 1 960 connectens: (R. Perrier, 1 898)
fusus (O.F. MUller, 1776) Zygothuria connectens R. Perrier, 1 898
=

?Thyone flexus Hodge, 1 867 expectans R. Perrier, 1 898

= Holo/huriafusus O.F. MOiler, 1 776 intestinalis (Ascanius, 1 805) Ostergren, 1 896
= Holo/huria papillosa AbiidglUlrd, 1 789 Fistularia mollis M. Sars, 1 835
= T. subvillosa Herouard, 1 890 = Holothuria inteslinalis Ascanius, 1 805
fusus subsp. mcdilerranea Madsen, 1 94 1 Thyonidium scabrum M. Sars, 1 868
gadeana R. Perrier, 1 898 laelea (Theel, 1 886)
T. wahrbergi Madsen, 1 94 1 ?Holothuria aspera Bell, 1 892
roscovila Herouard, 1 890 = Holothuria laclea Theel, 1 886
Lep/open/acla Zygolhuria laclea (Theel, 1 886)
=

elonga/a (DOben & Koren, 1 846) maroccana Perrier, 1902

?Cucumaria eucumis (Risso, 1 8??) murrayi var. grandipes Herouard
= Cucumaria elonga/a Dilben & Koren, 1 846 roulei (Koehler, 1 896)
Cueumaria pentacles Forbes, 1 84 1 non (Linnaeus, Holo/huria roulei Koehler, 1 896
1767) rugosa Herouard. 1 9 1 2
/ergestina (M. Sars, 1 857) trirad/ata Heding, 1 94 2
= Cucumaria incurvata Perrier, 1 902 verrilli (Theel. 1 886)
= Cucumaria lergeslina M. Sars, 1 857 Holothuria verrilli Theel. 1 886
= M. oklaknemus Sluiter, 1 90 1
Order Dactylochirotida
Meseres
Fllmily Ypsilolburiidae aeaudum Heding, 1 935
Echinocucumis ingolji (Heding, 1940)
hispida (Barrett, 1 857) = Plicaslichopus ingolji Heding, 1 942
= Cucumaria hispida (Barrett, 1 857) Paelopolides
= E. typica M. Sars, 1 859 atlantica Herouard, 1 902
= Eupyrgus hispidus Barrett, 1 857 grisea R. Perrier, 1 902
Ypsilo/huria Paroriza
bifen/aculata subsp. atlenuata E. Perrier, 1 886 pollens (Koehler, 1 896)
talisman; subsp. talismanl E. Perrier, 1 886 = Stichopus pallens Koehler, 1 896
= ?Echinocucumis hispida var. abyssalis Koehler, 1 896 prahoi Herouard, 1 902
Order Aspidochirotida Pseudostichopus
atlanticus R. Perrier, 1 898
Family Stichopodidae
depressus Herouard, 1 902
Eostichopus globigerinae Herouard, 1 902
regalis (Cuvier, 1 8 1 7) lapidus Herouard
= Holothuria columnae Cuvier, 1 8 1 7 marenzelleri Herouard, 1 902
Holo/huria regalis Cuvier, 1 8 1 7 oceultatus von Marenzeller, 1 893
Holothuria triquetra Delle Chiaje, [84 I
villosus Theel, 1 886
=

Sfichopus regalis (Cul/ier, 1 8 1 7)

ScolOthuria
=

Parastichopus
herringi Hansen, 1 978
tremulus: (Gunnerus, 1767) = ?Dendrothuria simi/is Koehler & Vaney. 1 905
= ?Stichopus griegi ()stergren, 1 896 Synallactes
Holo/huria ecalcarea M. Sars, 1 859 crucifera R. Perrier, 1 898
=Holo/huria elegans O.F. MOiler, 1781
Holothuria tremula Gunnerus, 1 767 Fllmily Holotbu riidlle
= Slichopus richardi Herouard, 1 896 Gastroturia
Stichopus tremulus (Gunnerus) limbata R. Perrier, 1 899
Holothuria
Family SynaUactidae
(Roweothuria) arguinensis Koehler & Vaney, 1 906
Bathyplotes dakorensis Panning, 1 9??
natans (M. Sars, 1 868) ()stergren, 1 896 forskali Delle Chiaje, 1 823
B. ass/mills Koehler & Vaney, 1 905 = H. catanensis Grube, 1 840
= B. helerostylides Heding, 1 942
= H. nigra Peach, 1 845
B. ostergreni Ohshima, 1 9 1 5 = Slichopus selenkae B arrois, 1 882
= B . pap/Ilosus Koehler & Vaney, 1 905
helleri von Marenzeller, 1 878
=B. palagia/us Fisher, 1 907 = H. affinls Heller, 1 868, non Brandl, 1 83 5
=Holothuria na/ans M. Sars, J 868 (Thymiosicya) impatiens: (Forskal, 1 775)

- 3 49 -
Costello, Emblow and White (eds), 2001

= FislUiaria impatiens Forskal, 1 775 violacea Theel, 1 879

= H. botellus Selenka, 1 867 = Cryodora spongiosa Theel, 1 879
H. impatiens subsp. bicolor Clark, 1 93 8 L. brongniarli E. Perrier, 1 886
= H. impatiens subsp. concolor Clark, 1 92 1 = L. jourdainii Pelit, 1 885
H. impatiens subsp . /utea Clark, 1 92 1 = L . wyvillelhomsoni Theel, 1 879
= H. impatiens subsp. pulchra Clark, 1 92 1 Pannyahia
= H. truncata Lampert, 1 885 glutinosa Herouard, 1 902
(I'aneyolhuria) lentiginosa von Marenzeller, 1 892 Family Psychropotidae
mammala Grube, 1 840 Benthodyles
(Halodeima) mexicana Ludwig, 1 875 janthina von Marenzeller, 1 893
(Platyperona) parvula (Selenka, 1 8??) = B. lingua R. Perrier, 1 896
polii Delle Chiaje, 1 823 typica Theel, 1 882
sanelori Delle Chiaje, 1 823 = B. glutinosa R. Perrier, 1 902
= H. farcimen Selenka, 1 867 Psyehropoles
(Holothuria) stellali Delle Chiaje, 1 823 depressa (Theel, 1 882)
(Holothuria) tubulosa Gmclin, 1 788
= Euphronides auricularia R. Perrier, 1 902
= H. cavolin;; DeUe Chiaje, 1 84 1 Euphronides comuta Verrill, 1 884
= H. columnae Delle Chiaje, 1 823, non Cuvier, 1 8 1 7 = Euphronides lalismani R. Perrier, 1 896
H. maxima Delle Chiaje, 1 823 = Euphronides violacea Perrier, 1 902
= H. pentagnae Delle Chiaje, 1 823 grimaldii Herouard, 1 896
Order Elasipodida longicauda Theel, 1 882
= Neclolhuria translucida B elyaev & Vinogradov, 1969
Family Elpidiidae
= P. buglossa R. Perrier, 1 896
Ach/yonice
= P. furcola R.
Perrier, 1 896
myriamae Gebruk, 1 997
semperiana Theel, 1 882
Amperima
= Euphronides anchora Herouard, 19 J 2
furcala (Herouard, 1 899)
= Euphronides kerhervei (Herouard, 1 896)
= Ko/gafurcala Herouard, 1 899
= P. kerhervei Herouard, 1 896
rosea (R. Perrier, 1 896)
= Periamma rosea R. Perrier, 1 896 Family Pelagolhuridae
Ellipinion Enypniastes
delagei (Herouard, 1 896) eximia Theel, 1 882
= Scoloplanes delagei Herouard, 1 896 = Pelagothuria bouvieri Herouard, 1 906
Elpidia = Planklothuria diaphana Gilchrist, 1 920
glacialis Theel, J 876 Order Apodida
echinala (R. Perrier, 1 896)
Family SynaptidBe
TUlela echinala R. Perrier, 1 896
Epilomapla
Trpa
abyssieola Danielssen & Koren, 1 878 roseola (Verrill, 1 874)
Leplosynapta roseola Verrill, 1 874
Kolga
Euapla
hyalina Danielssen & Koren, 1 879
Elpidia nana Theel, 1 879
lappa (J. MUlier, 1 850)
SYllapla lappa J. MUlier, 1 850
Peniagone
=

Labidop/ax
azorica von Marenzeller, 1 893
diaphana (Theel, 1 882) buskii (M'lntosh, 1 866)
= Seoloonassa translucida Herouard, 1 899
= Synapla busk;; M'Intosh, 1 866
gracilis (Ludwig, 1 894)
= Synapta lenera Norman, 1 865 (nomen nudum)
digilala (Montagu, 1 8 1 5)
islandica Deichmann, 1 930
porcella R. Perrier, 1900
= Hololhuria digilata Montagu, 1 8 1 5
L. (Oes/ergrenia) adrialiea Heding, 1 93 1
purpurea (Theel, 1 882)
digitala var. profundlcola Kemp, 1 905
= P. ferruginea Grieg, 1 9 2 1 .
media Ostergren, 1 905
= P. vexillum R . Perrier, 1900
morlenseni Heding, 1 9 3 1
w)'villii Theel, 1 882
L . (Oeslergrenia) morlenseni Heding, 1 93 1
Penifpidia
=

similimedia Gage, 1 985

IUdwigi (von Marenzeller, 1 893)
southwardorum Gage, 1 985
Kolga IUdwigi von Marenzeller, 1 893
Ihomsoni (Herapalh, 1 865)
Psychroplanes
= Synapta digilala var. Ihomsonii Herapath, 1 865
obsolela (Herouard,1 899)
Leplosynapla
= Herouard, 1 9 1 2
Kolga foliacea
= Kolga obsoleta Herouard, 1 899
bergensis (Ostergren, 1 905)
= Synapla bergensis Ostergren,
1 905
= Peniagonefoliaeea (Herouard, 1 9 1 2)
decaria (O stergren,1905)
= Peniagone obsolela (Herouard, \ 899)
= Synapla decaria Ostergren. 1 905
Family Deimatidae galiiennil (Herapath, 1 865)
Deima = Synapla gal/ienn;; Herapath, 1 865
validulII subsp. valMum Theel, 1 879 =Synapla sarniensis Herapalh. 1 865
= D. allanlicum Herouard, 1 898 inhaerens (O.F. MUlier, 1 776)
D. blake! Theel, 1 886 =Hololhuria inhaerens O.F. MUlier, 1 776
Oneirophanta =Synapla duvernaea de Quatrefages, I S??
mUlabilis Theel, 1 879 makrankyra (Ludwig, 1 898)
0. allernala R. Perrier, 1 900 =Synapfa makrankyra Ludwig, 1 898
Family Laetmogonidae minula (Becher, 1 906)
Benthogone =Synapla mimlla Becher, 1 906
rosea Koehler, 1 896 Prolankyra
=B. quadrilineala Heding, 1 940 abyssicola (Theel, J 886)
Laelmogone = Synapla abyssicola Theel, 1 886

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 European Register of Marine Species

Rhabdomolgus scaber LUtken, 1 857

ruber Keferstein, 1 863 Echino.mma hispidum Semper, 1 868
=

Synapfula Hedingia
reciproeans (Forskal, 1 775) albicans (Theel, 1 886) Deichrnann, 1 93 8
= Synapta reciproeans ForskI, 1 775 Molpadia albicans auett.
=

Family Chiridotidae Trochos/oma albieans Theel, 1 886

Achiridota mediterranea (Bartolini Baldelli, 1 9 1 4 ) Tortonese,
profunda Reding, 1 935 1965
Chiridota Trochostoma mediterraneum Bartolini Baldelli, 1 9 1 4
abyssicola von Marenze"er, 1 893
laevis (0. Fabricius, 1 780)
C. groenlandiea Heding, 1 935
C. spirourna Heding, 1 935
= Hololhuria laevis O. Fabricius, 1 780
Hololhuria pellucida Vahl, 1 806
Trochodola
furcipraedila Sa1vini Plawen, 1 972
venusta (Semon, 1 887)
= Chiridola venusla Semon, 1 887
Family Myriotrocbidac
Acanfhofrochus
mirabilis Danielssen & Koren, 1 879
Myriotrochus
balhybius H.L. Clark, 1 92 0
clarki Gage & Billett, 1 986
eurycyclus Heding, 1 93 5
geminiradialus Sa1vini-Plawen, 1 977
giganteus Clark, 1920
rinkii Steenstrup, 1 85 1
Iheeli Ostergren, 1 905
vilreu8 (M. Sars, 1 872)
= Chiridofa brevis Huxley, 1 852
= Chiridola laevis C.GJ. Petersen, 1 898
= Oligolrochus vitreus M . Sars, 1 872
Parvolrochu8
belyaevi Gage & B illett, 1 986
Siniofrochus
myriodontus Gage & B illett, 1 986
Trochoderma
elegans Theel, 1 877
Order Molpadiida
Family Molpadiidae
Cherbonniera
ulriculus Sibuet, 1 974
Molpadia
blakei (Theel, 1 886)
=Paralrochosloma spiniferum Heding, 1 932
=Trochostoma blakei var. excenlrica Herouard
Trochos/oma blakei Theel, 1 886
=Trochosloma grossularia Herouard
borealis M. Sars, 1 859
=Ankyroderma affine Danielssen & Koren, 1 882
=Ankyrodermajeffreysii Danielssen & Koren, 1 87 8
Haplodaetyla arc/iea v o n Marenzeller, 1 877
=M (Paramofpadia) dipfoa Heding, 1 935
=M oolitica Mortensen, 1 9 1 4 non Pourtali:s, 1 85 1
Troehos/oma arc/iea Danie1ssen & Koren, 1 880 non
(von Marenzeller, 1 877)
= Trochostoma boreale (M. Sars)
Trochosloma Thomsonii Danielssen & Koren, 1 879
foricala (R. Perrier, 1 898)
= Ankyroderma forica/urn Perrier, 1 898
maroccana (Perrier, 1 898)
= Ankyroderma rnaroceana Perrier, 1 898
musculus Risso, 1 826
= Ankyroderrna danielsseni Theel, 1 886
= Ankyroderma hispanicum Petit
Ankyroderma perrieri Petit
= Eumolpadia asaphes Heding, 1 935
Haplodaclyla mediterranea Grube, 1 840
= Trochostoma violaceum Theel, 1 886
ooli/ica (Pourtales, 1 SS I )
Family Eupyrgidae
Eupyrgus

- 35 1 -