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‘Chapter 2 STRUCTURAL 'S OF THE RAIN FOREST. P.8, TOMLINSON INTRODUCTION: method of climbing, or the epiphytes according to their water economy, or the herbs according to their method of vegetative spread. ural an d to diffe ses Of tropical rain forest may levels of resolution. This ac inly with the major tropical rain Is, Le; trees, and atlempistoestab- ARCHITECTURE: compo ¢ immediately obvious processes involved in Following Hallé and Oldeman (1970) and Hallé lopment, An analysis of the architecttire al, (1978) it is recognized that each individual dividual trees provides the analytical too! for species has a precise and genetically determined Reassessment of plant thterdetion. This assessment growth plan or architectural model, while the ex gross morphological diversity is a fesumé of the pression of this growth plan at any. one develop= h adopted by Hallé ét al (1978) Detailed PENG stage may be said to represent the wrchi- phological accounts of the biological con- tecture of the plant. These concepts differ from of rain forest plants are found in Schnell (you implied in the general terms life-form or ‘ore (1975), Longman and Jenik growth habit in that the latter are both, ee bove-ground parts are considered concepts and refer to the end-product — e.s y detail. The architecture and interaction of shrab, soe stems in the rain forest have been sum- 14 ogni marized re common lite-form: (1) Canopy trees (including Richards, 1957) (2) Understorey trees ) Shrubs a (4 Lianes or woody climb stranglers) (5) Unierstorey herbs (6) Vascular epiphytes CO) Vaseular parasi (8) Vascular sapropl Each of these may be eaps subdivision, e.g, ‘the’ I P.B. TOMLINSON gp contrast, in natural environments disturbance. TT jay not conform to its architectur dhe tree may gn tbe exten (0 wien (He Pina depend Sful cireumstances, as BY StOrm or yee or by crown interaction. Fallow. 1974) we hrave to add the concept of srowth response to environmental grow analysis of tree architecture, as when broken as when existing be traumatic, stored, of adaptiv 4, The normal reiterative re- reorientated ms, which are not ex- |, to develop and in chitecture of lest and most ie substitution of a ier by a latent , but the kinds ;compassed in the concept of nd yaried. In many conifers ly part of the original is repeated as, for example. the plagtotropic ‘ches in Araucaria (Veillon, 1978), The reiter- ative ability of a tree is probably as important an ecological parameter as architecture itself, if not greater, but varies enormously. At one end of the scale there is no possibility of reiteration, as with

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