Anthropology
163
Midterm
1
03/22/09
R.M.
La
Jeunesse
The
following
is
the
definition
of
a
species
used
for
this
paper.
Lets
say
that
we
have
a
species
(S)
and
beings
that
are
of
that
species
and
can
reproduce
with
each
other
(s).
Each
s
in
S
has
a
certain
set
of
traits/genes
that
they
have
in
common,
if
a
being
does
not
have
that
certain
set
of
traits
in
common
with
S,
then
that
being
will
not
be
able
to
reproduce
with
other
s
or
is
not
a
member
of
S.
Let
the
set
of
traits/genes
that
are
found
is
S
be
{a,b,c,
X}.
Where
a
=
a
set
of
physical
reproductive
traits/genes
b
=
a
set
of
behavioral
traits/genes
affecting
reproduction
c=
a
set
of
other
traits/genes
specific
to
S,
or
affect
reproduction
with
an
s.
X
=
a
set
of
traits/genes
that
do
not
affect
reproduction.
s
is
said
to
be
a
member
of
S
and
can
reproduce
with
other
members
of
S,
if
s
contains
the
set
{a,b,c}
If
s1
=
{a,b,c,A}
and
s2={a,b,c,B}
since
s1
and
s2
contains
the
set
{a,b,c},
then
they
are
members
of
S,
and
can
reproduce.
If
s1
=
{b,c,
A}
and
s2={a,b,c,
B}
since
s1
does
not
contain
{a},
then
s1
is
unlikely
to
be
able
to
reproduce
with
s2.
If
s1
=
{a,c,A}
and
s2=
{a,b,c,B}
since
s1
does
not
contain
{b},
then
s1
may
be
physically
capable
of
reproducing
with
s2,
but
s1
may
not
attempt
to
reproduce
with
s2,
or
s2
or
s3
may
not
let
s1
reproduce
with
s2.
If
s1
=
{a,b,A}
and
s2=
{a,b,c,B}
since
s1
does
not
contain
{c},
then
there
is
some
trait
that
is
missing
or
present
in
s1,
that
is
hindering/preventing
s1
from
reproducing.
[Question
1]
Macroevolution
is
the
process
of
an
S
changing
over
time,
or
the
split
of
an
S
into S1 and S2. There are two competing views that exist when explaining the
process of macroevolution; punctuated equilibrium theory and the gradual
development
theory.
The
punctuated
equilibrium
theory’s
basic
premise
is
that
macroevolution
occurs quickly (evolutionarily speaking) after a certain event takes place. The event
results in some sort of barrier (roads, rivers, mountains, islands, latitude, resource
location, etc.) that separates S1 and S2. At the time of separation the set of
traits/genes in S1 and S2 are the same. Over time, S1 and S2 adjust (quickly) to their
different environments, and the traits/genes {a,b,c} may change in S2, but not in S1,
and once those traits differ, then we could state that S1 and S2 are two different
species. If the environment remains constant, then {a,b,c} has nothing acting on it to
change. This would imply that if no barrier separates S1 and S2, then the traits
{a,b,c} will not change, and thus S will remain constant.
The gradual theory of macroevolution contrast with punctuated equilibrium
theory with respect to the clause: if no barrier separates S1 and S2, then the traits
{a,b,c} will not change, and thus S will remain constant. Gradual theory implies the
idea of genetic drift, or that over time, the traits/genes {a,b,c} in S could change
without any physical barrier, only time and distance is needed. The classic example
has to do with the lizard reproduction problem. S1 can reproduce with S2, S2 can
reproduce with S3, S3 can reproduce with Sn, however S1 cannot reproduce with
Sn. This problem illustrates that speciation is a continuum, not a “punctuation”.
(This creates a problem with the definition stated above, lets correct it).
Definition
for
genetic
drift
causing
speciation
{a,b,c}
are
a
set
of
traits/genes
that
must
exist
for
s
to
reproduce
with
a
member
of
S,
for
example,
if
a
=
{w,x,y,z}.
Assumption:
An
s
can
have
trait/genes
{a}
if
s
has
the
majority
of
the
elements
of
a:{w,x,y,z}.
So,
s1
can
physically
reproduce
with
s2,
if
a1
=
{w,x,y,z}
and
a2
={j,x,y,z}
s2
can
physically
reproduce
with
s3,
if
a2
=
{j,x,y,z}
and
a3
=
{j,k,y,z}
s3
can
physically
reproduce
with
sn,
if
a3
=
{
j,k,y,z
}
and
an
=
{
j,k,l,z
}
Therefore,
Sn
cannot
physically
reproduce
with
S1,
because
an
=
{
j,k,l,z
}
and
a1
=
{w,x,y,z}
implying
that
Sn’s
{a}
is
not
found
in
s1,
and
if
Sn’s
{a}
is
not
found
in
s1,
then
S1
cannot
physical
reproduce
with
Sn.
This
pattern
could
also
be
true
with
traits/gene
sets
{b,c}
However,
the
distinction
between
gradual
and
punctuated
theories
seems
to
be misconceived, because it appears they could be identical. The difference between
the two theories is the notion of time and the definition of “barrier”. The barriers
that the punctuated theory shows, still results in genetic drift, the difference is that
barriers create two paths for genetic drift to occur. Because if you have S, and then a
barrier creates S1 and S2, initially S1 = S2, and over time in the different
environments genetic drift will occur, and later S1≠ S2. This should happen quicker,
because S1 and S2 can no longer interact. The drift could happen even quicker, if
genetic drift was already occurring, and a barrier developed between S1 and Sn.
Gradual evolution theory also implies a barrier; time and distance. The
barrier of time implies that a certain environment will not always be the same. Over
the course of a million years, there can be significant change in an environment
(weather, resource location, etc.), which would imply the traits/genes {a,b,c} of the
species might need to change to adapt to the change of the environment, or the
species would become extinct. With distance, as the species spreads out, the
environment the species lives in changes. The species spectrum (S1~Sn) might not
be
separated
by
a
physical
barrier,
but
s1
and
sn
are
separated
by
a
barrier
of
distance,
because
s1
is
unlikely
to
travel
500miles
to
reproduce
with
sn.
Also,
time
could be look at as a barrier of distance.
However, the issue that seems be in debate by the two theories is the
assumption made in the definition of genetic drift. An s can have trait/genes {a} if s has the
majority of the elements {w,x,y,z}. A punctualist might state that the assumption is false,
and claim that every element in {w,x,y,z} needs to be present for reproduction to
occur, and then argue what does “majority of elements” constitute.
The area of empirical evidence that is being used to debate this problem is
the fossil record. Punctualist state that because the fossil record is not continuous,
then that is clear evidence that evolution happens punctually. However, just because
there is an absence of evidence in the fossil, does not imply that evidence does not
exist. With this in mind, gradualist would state that the fossil record is a snap shot of
the evolution of a species, and the missing gaps in the fossil record have not been
found, or since fossilization is a rare occurrence, then a fossil might not have been
made of that stage in evolution.
In humans the fossil record from about 2million years ago seems
inconclusive about how humans evolved because there are missing links. The fossil
record suggest that modern homo sapiens originated on the East Coast of Southern
to Northern Africa mostly along the Nile river, however it is unclear if there was
gradual evolution or an event that separated homo sapiens from the other primates.
However, if we strictly go by the evidence in the fossil record, it would suggest that
man came from punctuated events.
Some
would
say
that
the
fossil
record
is
limited
in
its
ability
to
explain
human evolution, and we need to look at other sources to find the answers. Now
how else could we look at speciation in Humans? (Question 4) Before we can
answer that question we need to review how speciation occurs in mammals and
then take a look back at humans. Speciation of mammals would occur when the
elements that govern reproduction {a,b,c} are no longer congruent between two
populations of mammals that had a common ancestor.
Minimum
requirements
for
speciation
There
is
an
S
which
has
traits/genes
{a,b,c,
X}.
A.
If
a
barrier
comes
to
exist
in
S,
then
it
will
result
in
S1
and
S2.
B.
If
the
barrier
prevent
S1
and
S2
from
interacting,
then
the
gene
pools
in
S1
and
S2
to
freely
vary
from
each
other.
C.
S2
is
said
to
speciate
when
traits/genes
{a,b,c}
differ
from
those
is
S1.
The
most
crucial
elements
in
all
species
including
mammals
are
the
traits/genes that are directly responsible for reproduction, {a}. In mammals these
are traits such as sexual organs, creation of sex cells, antigens, incubation of the
fetis, incubation time, route of exit of the fetis, mammary glands, health of the
mother, 2 sexes, and chromosome number to be a few examples. Within a species
these traits should in the most part be similar, however any significant deviation in
these traits, may make reproduction impossible. If the trait change hinders
reproduction, then it would have a very low chance of changing across species.
Implying that trait/gene {a} should remain fairly constant across species of
mammals. The only problem here is that chromosome number can changes across
species.
Not
sure
how
to
explain
that.
The
behavior
component
of
mammalian
species
{b}
should
vary
much
more
in mammals than the reproductive traits. Behaviors that aid in reproduction in
mammals are selective breeding, socialization, mother and fathering, migration,
imitation, altering environment, etc. Not all of these traits are found in all mammals.
A behavior that directly relates to mammalian reproduction is mate selection. If a
set of females in S will only reproduce with a male of S who does behavior X, and
there is another set of females in S that will only reproduce with a male who does
behavior Y, then this creates a behavior barrier, S1=S2. Then in time, genetic drift
and selection on the favored behavior might result in two distinct species.
The {c} component of a species has to do with traits/genes that are indirectly
related to reproduction (these can be physical traits or behaviors). These are traits
involving environmental adaptations, disease resistance, resource acquisition
ability, size, basically anything that can be selected for or against. In mammals one
of these traits might be resource preference. If S1 prefers berries on bushes, and S2
who is smaller than S2 develops a preference for berries in a tree, then in time these
groups might speciate.
However, where can all of this variation come from? That is the beauty of the
{X} component. Since natural selection is not acting on {X}, then the elements in {X}
might cause a trait to emerge that is beneficial. Once that beneficial trait emerges
and gives the individual an advantage, then it could either spread to the S, or result
in speciation. Could it be possible that the extra chromosome problem be a member
of
{X}?
Human
variation
should
follow
a
similar
path
to
mammals,
if
there
is
any
change in the {a,b,c}’s then speciation might occur. The most significant punctuation
that could cause an S1 and S2 in early man may have been our tendency to form
social groups. If each social group tended not to interact (which is evident with
outsider hatred problems), then the social group would be a barrier to early
humans. Between the groups, there would selective pressure for resources to be the
dominant group, and it could be argued that as the communication in a group
increases, then the strength of the group increases. If true, then this could provide a
nice environment for humans to emerge from.
A major problem that occurs in humans is that despite being spread across
the entire globe for 60,000years, we did not speciate. If rapid speciation does occur
as punctuality claims, then why could Pocahontas and John Smith have child? Is
there something special about humans that prevent us from speciating?
[Question 2]
One of the first genetic systems discovered is ABH series in the human blood
system. These are a series of traits that generally fall into the variability case {X} of
the {a,b,c,X} distinction. The ABH series generally does not affect ones ability to
reproduce. However, there have been times in recent human history that this blood
group series allowed some individuals to reproduce and others not be able to
reproduce, making the blood groups a {c} at times. For example, there have been
epidemics where a certain blood group was resistant to pathogens.
The blood groups ABO are composed of two components, creating 4 distinct
groups;
A,B,AB,O.
The
first
component
is
the
structure
on
the
outside
of
the
blood
cell,
called
antigens.
These
antigens
allow
for
the
immune
system
to
recognize
and
not attack the blood cell. If the wrong antigen type is put in a person, then the
immune system will identify the blood as an invader, and destroy it. Blood type A,
has antigen‐a, B has antigen‐b, AB has antigen‐a and antigen‐b, and O does not have
any antigens.
The second component is a set of anti‐bodies that would detect the presence of
the other blood types. Blood group A would have antibody‐B, B would have
antibody‐A, AB would not have any, and O would have antibody A and B. Then to
further complicate the blood system, there is a another set of antigens the Rhesus
group, which is classified as + or ‐, depending on the presence of certain R‐antigens.
When transferring blood, one has to be careful. If B blood was placed into a
person with A blood, then the person’s antibodies‐B would reject the B blood. Since
O blood has anti‐bodies for A and B blood, then an O blooded person can only take O
blood, but since the blood has no antigens, then it can be donated to everyone. AB
blood on the other hand, can receive blood from everyone because there are no
antibodies, but cannot donate to other groups, because the other person’s
antibodies would connect to the AB bloods antigens.
The disease that caused the black plague was delivered by vector of a flea,
and then transferred from human to human when it became pneumonic. This
particular disorder seems to be more successful in the people with O blood groups.
It is possible that the O blood would be easier to attach to than the A and B blood,
because O blood has no antigens. Since, the plague has a high mortality rate,
combined
with
the
success
it
had
in
the
O
blood
system,
then
areas
where
the
plague
occurred
have
a
lower
population
of
O
blood
groups.
This
would
be
why
the
O
blood
is less common in Europe, than other regions of the world.
The same problem occurred with small pox. Small pox has an antigen system
similar to antigen‐A. This would mean that the A‐blood system would identify small
box as if it is blood, and not get rid of it. However, since B and O blood have
antibodies‐a, then these systems would flush small pox out of the body. Thus, those
with the A blood system would more likely die from small pox, thus blood type A
would be less prevalent in areas where there was a small pox epidemic occured.
Since small pox has been very common in India, this would explain why the A‐blood
group is less common in India.
It could be predicted that due to recent human activity that the distribution
of blood types will slowly become evenly distributed. The reason why is because
cultural practices. Europeans created the environment that caused the black plague,
by creating the sanitary conditions that favored the rat that carried the flea that
possessed the plague. Europeans have since learned from that experience, and now
practice sanitation better. Also, humans have created a way to combat the plague
through antibiotics, and small pox through a vaccine. Thus, in the future these
pathogens will no longer have the selective pressure they once had, and blood
groups should back fall back into free variations.
Questions
3.
Humans
have
many
physical
traits
that
can
fall
into
the
{X}
category
stated
above, but on occasion some traits can become a member of {c}. These traits are
often
explained
in
terms
of
dominant
and
recessive
genes,
represented
as
Aa.
If
the
dominant
trait
is
present,
then
that
A
will
cause
the
phenotype.
If
only
aa,
then
the
recessive phenotype will emerge. However, there could be multiple systems acting
on one phenotype, polygentic traits, represented as Aa, Ba, and Ca. Better yet, some
phenotypes could be interacting with other trait systems, or relying on other
systems to be activated.
A good example is the skin tone in humans. In the past, humans found
themself in an environment that favored thinned hair, and then the need for
pigmentation in the skin. The initial environment was in the tropics near the
equator, where there is a high degree of UV exposure. UV light from the sun can
damage and cause the bonds in protein and DNA to break down, and cause skin
diseases. So, those that have better protection from UV light, will live longer and be
more active during the day, than those that have less protection. Those that were
heavily pigmented had an evolutionary advantage over those who were not.
The exact opposite occurred when humans migrated northward. As one
moves away from the equator, the amount of UV light present decreases. That would
be no problem if UV light was not needed for vitamin D production, however it is.
This would explain why as one heads north of the equator, pigmentation in humans
decrease. Because vitamin D is needed for a child to develop bones, and if the
vitamin production is hindered, then the child might not live to adulthood.
Hair and eye color seem to correlated with a person’s pigmentation. When a
person has very dark pigmentation in their skin, their hair is very day and their eye
color is almost black. However, when a person has little pigmentation in their skin,
then
there
hair
is
blonde
or
red
(low
pigmentation)
and
eye
color
is
blue
(low
pigmentation).
This
suggests
that
the
pigmentation
of
skin,
hair,
and
eye
color
all
depend on similar melanin regulating genes. If those genes malfunction, then we
have albinism, affecting skin, hair, and eyes.
However, this does not fully explain hair color, because we have blonde hair
and red hair when melanin is low. This suggests that hair color could have two
systems regulating the color. Melanin is the dominant color, and covers up the red
coloring in heavy pigmented people. The same is true with eye color, if a person has
low pigment, then they can have blue or green eyes. This would suggest that eye
color has other systems regulating color as well.
[Question 5.]
Anthropologist tends to view stature as being a genetic trait that is free to
vary, {X}. Mostly, the pituitary gland regulates stature. The average height for
humans is 5’5ft. There are genetic factors that regulate height, but the
environmental factors seem to have a significant affect, which could imply that the
distribution of stature has more to do with culture, than genetic drift. The general
consensus is that stature is increasing across populations.
Simple
way
of
determining
the
affect
of
culture
on
stature.
Let
M
=
the
elements
that
that
affect
stature.
Where
0
≤
M
≤
n
n
=
maxium
amount
of
resources
that
can
affect
stature.
P
=
Population
#.
D
=
the
distribution
of
the
resource
M
in
P
If
M=x,
then
stature(S)
=
g+(0.5)x,
n
=
6
g
=
genetic
base
height
Assume:
g
=
3.0ft
In
society
A,
MTotal
=
25,
P
=
5,
and
D
=
evenly
distributed.
M1
=
5
S1=5’5,
M2=5
S2=5’5,
M3=5
S3=5,
M4=5
S4=5’5,
M5=5
S5=5’5
Save
=
5’5.
In
society
B,
MTotal
=
25,
n=6,
P
=
5,
and
D
=
60%(Pa=2)
and
40%(Pb=3).
M(Pa)
=
15
Ma1
=
7.5
Sa1=
6.0ft,
Ma2
=
7.5
Sa2
=
6.0ft
M(Pb)
=
10
Mb1
=
3.33
Sb1
=
4.6ft,
Mb2
=
3.33
Sb2
=
4.6ft,
Mb1
=
3.33
Sb1
=
4.6ft
STotal
=
2(6.0ft)
+
3
(4.6)
ft
=
25.8,
SAve=
5.15ft
Even
though
M
is
equal
in
population
A
and
B,
the
distribution
of
resource
can
affect
average
height.
Culture
affects
stature
by
either
increasing
or
decreasing
the
elements
of
M.
For instance, a culture that invented the boat is going to be able increase its supply
of protein to the population, increasing the value of M, which would result in high
stature size. However, a society could have a class that holds the fish for themselves,
and does not let the majority of the population access the protein. This would be an
example of the value of M being high, but D being offset. This would create a class of
tall people, and short people.
Initially as a population increases, the resources(M) would have to be
distributed to the increasing population. If D remains constant, then the value of M
for each person would decrease, and thus their stature would decrease. However
modern society has created environments that provide a surplus of the resources
that affect stature, and thus M is almost at n for most members of the society, thus it
currently appears that as population increases, stature increases.
I assume that affective population size implies a society that is self‐
sustaining. The affective population for maximum stature would be for M=n for
every member of the population. On an island there are a limited amount of
resources available, and thus the value of M is not is not going to increase as the
population increases. Therefore, as the population on an island increases, the values
of M remains constants, and D remains constant, then the stature of the population
will
have
to
decrease.
If
D
varies,
then
only
the
deprived
portion
of
the
population
will
shrink,
or
die
of
malnutrition.
This
shrink
in
stature
could
account
for
dwarfism
in areas that are able to sustain a population, if the resource of M is low, or hindered
by a cultural practice.
Island
example
M
=
20,
D
=
even
Initial
population
P
=
4
Mn
=
M/P
=
20/4
=
5
If
Mn=5
Then,
Sn
=
5.5ft
SAVE
=
5.5ft
Future
population
P=8
Mn
=
M/P
=
20/8
=
2.5
If
Mn
=
2.5
Then,
Sn
=
4.25ft
SAve=4.25ft