Kelompok 1/Pend.

Biologi 6B/Vertebrata
1. Khoirunnisa

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2. Nuramelia

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3. Nuri Purnama 109016100060

4. Syarifah Aeni 109016100062
5. Dwi Nanda B P

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6. Hadi Suari

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Kelas : PISCES
Ciri-ciri Umum:
1. Berdarah dingin (poikiloterm).
2. Bernafas dengan insang
3. Permukaan tubuhnya licin, bersisik, dan pergerakannya menggunakan sirip.
4. Habitatnya ada yang di air tawar, air laut, dan air payau.
5. Merupakan kelompok vertebrata (bertulang belakang) yang paling beraneka ragam
dengan jumlah spesies lebih dari 27.000 di seluruh dunia.
6. Hanya memiliki telinga bagian dalam.
7. Peredaran darah tunggal.
8. Memiliki otak yang dibungkus oleh kranium yang berupa tulang rawan.
9. Memiliki sepasang mata.
10. Memiliki organ keseimbangan yang disebut gurat sisi (linea lateralis).
Ciri-ciri Khusus
1. Kelas Agnatha atau Cyclostomata (Ikan tidak berahang)

Tubuh seperti belut

Kulitnya lunak berlendir

Tidak ada pasangan sirip, beberapa jenis memiliki sirip ekor dan sirip
punggung, contoh belut laut.

Lubang hidung tunggal di sebelah dorsal

Faring dengan tujuh atau lebih celah insang dan terbuka secara tidak langsung

Noctocorda persisten (tetap ada seumur hidup) dan secara tidak sempurna
dilengkapi dengan vertebrae cartilago

Otak terdiri dari lima bagian : telensefalon, diensefalon, mesensefalon,

metensefalon, dan mielensefalon.

Ada 10 batang saraf kranial.

Ada yang mempunyai proneros (ikan hantu) dan mesonefros (belut laut)

Tidak ada sistem porta ginjal (angkutan atau aliran ginjal).1

Kelamin terpisah (hermafrodit).

Hidup dari cari cairan tubuh atau organ tubuh ikan lain dengan jalan
mengisapnya dengan mulutnya yang bulat. Ikan membuat lubang pada tubuh
korbannya dengan memarut tubuh ikan korban menggunakan mulutnya.

Siklostomata hidup dalam air tawar dan air laut.2

Badannya memanjang berbentuk silinder sedangkan ekornya pipih . Kulitnya

licin tanpa sisik , dilengkapi kelenjar lendir (mucus ) . Siri tengah dorsal
disokong oleh tulang-tulang sirip (tulang rawan ) . Matanya ada sepasang

Mulutnya ventro anterior dan merupakan mulut pengisap , dipingiran terdapat

tentakel . Kantong hidung terdapat disebelah tengah atas dan jumlahnya hanya
satu

Tengkorak kepala dan lengkung insang (viceral ) terdiri dari tulang rawan dan
notochord masih didapati dilengkapi archus neuralis yang tidak sempurna

Jantung terdiri dari dua ruang ( serambi dan bilik ) . Darah merah berbentuk
bulat-bulat dan berinti juga memiliki butir-butir darah putih serta jantungya
hanya memiliki satu ventrikel.

Insang terdiri dari 6 sampai 14 pasang terdapat di sisi pharynx berbentuk

kantong .

Ginjalnya sepasang bermuara di papil urogenitalis.

Temperatur tubuhnya tridak tetap (poikilothermus).

Alat kelamin atau (gonad ) sebuah tidak memiliki saluran kepapilla urogenitalis
. Pembuahan terjadi di luar tubuh . Telur yang sudah dibuahi menetas menjadi
larva (=ammocoete = pride ) dan ada yang langsung menjadi hewan (anak ) dewasa.

Otaknya berkembang naik , dengan 8 atau 10 pasang saraf cranial .

mempunyai alat pendengar dengan 1 atau 2 bentuk saluran setengah lingkaran

1 Mukayat Djarubito Brotowidjoyo, Zoologi Dasar, Erlangga, Jakarta, 1994, hal. 181.
2 Ibid, hal. 182.

dan mempunyai indra pembau .

hidup diperairan tawar dan laut

Air ludahnya mengandung bahan kimia yang mampu mencegah pembekuan

darah(ikan Lamprey)

Hidup secara parasit ataupun pemakan bangkai (Scavanger )

gigi dari zat tanduk, dan mempunyai lidah

siripnya tidak berpasangan

.rangka tubuhnya tersusun dari tulang rawan.

2. Kelas Chondrichthyes (Chondros = tulang rawan, ichthyes = ikan).

Ikan dengan mulut ventral disokong oleh rahang

Skeleton dari tulang rawan

Kulit tertutup dengan sisik-sisik plakoid (yang berasal dari kombinasi

mesoderm dan ektoderm)

Ada dua pasang sirip, sirip kaudal kebanyakan heteroserkal (lobus dorsal lebih
besar)

Ruang hidung berpasangan.

Faring dengan 5-7 celah insang (pada chimaera faring tertutup oleh operkulum
tunggal).

Sebagian notochorda diganti oleh vertebrata yang lengkap

Otak terbagi menjadi 5 bagian, dengan 10 saraf kranial

Pada ikan dewasa terdapat mesonefros

Ginjal terdapat sistem porta

Dalam usus terdapat katub-katub spiral

Kelamin terpisah, Fertilisasi eksternal atau internal

Ovipar atau ovovivipar

Secara praktis semua hidup di laut, contohnya ikan hiu, ikan pari, chimaera.3

3 Ibid, hal. 185.

3. Kelas Osteichthyes

Tulangnya tersusun dari senyawa osteon

mulut dan lubang hidungnya ventral, celah-celah pharyngeal tertutup (tidak

terlihat dari luar)

jantungnya hanya memiliki satu ventrikel. Jantung beruang dua, darah

berwarna pucat, mengandung eritrosit yang berinti dan leukosit.

Ikan ini juga mempunyai sistem limfa dan sistem porta renalis.

Mempunyai hati yang berkantong empedu.

Lambung dipisahkan dari usus oleh sebuah katup, mempunyai kloaka, tetapi
tidak jelas adanya pankreas.

Terdapat gelembung renang. Mempunyai gurat sisi, indra mata, telinga dalam
dengan tiga saluran semisirkulerdan memiliki otolit untuk keseimbangan.

Bernapas dengan insang yang memiliki tutup insang (operkulum).

Ukuran 1-600cm

Sebagian besar, tubuhnya tertutup sisik, memiliki kelenjar mucus

Pada tahap embrio memiliki 6 pasang insang, setelah dewasa tinggal 2-5 insang

Sirip vebtral dan dorsal hamper sama panjang

Ususnya panjang, fertilisasi eksternal, telur bercangkang lunak, dan hidup di air
tawar, payau, dan asin

Bachtiar
Ifftahussadiyah
Raden Mas Antonio
Reni Destriyani
Ria Mahardika
Syifa fauziah

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Ikan secara garis besar dibagi menjadi dua kelompok, yaitu Agnatha dan Gnathostoma.
1. Agnatha
Agnatha merupakan kelompok ikan yang pertama kali dikelompokkan oleh Cope (1889)
bersama dengan Gnathostoma. Agnatha merupakan kelompok ikan tidak berahang. Ada
tiga macam kelompok besar Agnatha, yaitu:
a.Osteostracida
Memiliki ciri-ciri berupa kepala yang memiliki perisai, sepasang bukaan
(diperkirakan merupakan organ mata), dan bukaan tunggal di bawah kanal naso-

hipofaringeal.
b.Anaspida
Memiliki tubuh yang streamline (pipih) dan sisik yang kecil berguna untuk menjadi
perenang yang tangguh. Ekor hiposersal mungkin dihasilkan dari bagian anterior
(berenang). Di bagian belakang kepala ada barisan pori-pori yang dipertimbangkan
sebagai insang.
c.Pteraspidomorphida
Pteraspidomorphida merupakan kelompok Agnatha dengan kepala dan tubuh yang
dilindungi oleh lapisan yang keras. Pteraspidomorphida merupakan kelompok
Agnatha yang paling sukses secara taksonomi dan jangka waktu dalam penemuan
fosil.
Ada tiga kelompok utama ikan yang masuk dalam kelompok Agnatha, terutama
yang masih hidup. Adapun ketiga kelompok tersebut adalah slime hag, Hag fish, dan
Lamprei.
Adapun kelas Agnata ini Dibagi menjadi 2 ordo, yaitu:

Ordo : Mixiniformes, contohnya hagfish (Myxine sp).

Ordo : Cephalasipidomorphi, contohnya lamprey (Pretromyzon sp.)

2. Gnathostoma
Gnathostoma dikarakterisasi dengan adanya rahang, atau sering dikenal dengan ikan
berahang. Ada 4 kelompok (kelas) utama dari Gnathostoma, yaitu Chondrichtyes
(kelompok ikan yang rangka tubuhnya tersusun dari tulang rawan) dan Osteichtyes
(Teleostei) dan dua kelompok yang sudah punah, yaitu Acanthodii serta Placodermi.
Kelas Chondrichtyes dibagi menjadi:

Ordo : Rajiformes (Hipotremata), contohnya ikan pari

Ordo : Squaliformes (Pleurotremata), contohnya ikan hiu

Ordo : Chimaeriformes (Rattfish), contohnya ikan chimaera

dua subklass utama, yaitu Elasmobranchii (hiu dan ikan pari) dan Holocephalii (chimaeras).
Sementara itu, kelompok Osteichtyes dibagi menjadi:
1.) Kelas Osteichtyes ; kelompok ikan yang rangkanya disusun dari tulang sejati. Dibagi
menjadi beberapa ordo, yaitu :

Ordo : Lophiiformers, contohnya ikan piranha

Ordo : Echeneiformers, contohnya ikan remora

Ordo : Perciformers, contohnya ikan kakap, belosoh, gerik, kapasan,

rejung
-

Sub Ordo : Trichiuroidei

Sub Ordo : Scombroidei

Sub Ordo : Stromateoidei

Sub Ordo : Acanthuroidei

Sub Ordo : Blennioidei

Ordo : Clupeiformers, contohnya ikan Bandeng

Ordo : Coelacanthiformers, contohnya Latimeria chalumnae

Ordo : Beloniformers, contohnya ikan terbang

Ordo : Siluriformers, contohnya ikan lele

Ordo : Mugiliformes, contohnya ikan kuniran, ikan belanak lancip

Ordo : Saccopharyngiformers, contohnya belut pelikan

1. Fosil Gnathostoma
Fosil Gnathostoma dibagi menjadi dua kelompok utama, yaitu Acanthodii dan
Placodermi.
a. Acanthodii
Kelompok ini sering dijuluki dengan hiu berduri. Secara relatif memiliki bentuk
yang memang mirip dengan ikan. Hidup pada pertengahan periode Silurian dan
punah pada pertengahan Permian (290-245 Miliar tahun lalu). Kebanyakan
kelompok Acanthodii merupakan kelompok ikan kecil yang tidak lebih dari 20 cm,
tetapi ada yang ukurannya diperkirakan lebih dari dua meter. Acanthodii memiliki
duri yang kuar sebelum sirip dorsal, sirip anal, dan sepasang sirip. Memiliki sisik
yang mirip dengan mahkota.
Acanthodii memiliki karakter berupa mata yang lebar, diperkirakan merupakan
perenang pada perairan tengah, beberapa di antara mereka mengembangkan gigi
yang kuat, sedang sebagian yang lain merupakan filter feeder, yaitu memeroleh
makanan dengan menyaring air yang ada disekitarnya.
b. Placodermi
Placodermi merupakan sekelompok yang sangat variatif dengan lapisan pelindung
tubuh yang sangat keras dan tebal. Kelompok ini berkembang pada zaman
Devonian (408-360 miliar tahun lalu). Kelompok ini memiliki rahang yang sangat
primitif.

Placodermi merupakan kelompok dengan yang pipih dorsoventral. Semua memiliki

pelindung pada kepala yang mampu digerakkan dengan pelindung pada bagian
depan dari tubuhnya.
c. Actinopterygii
Asal muasal dari kelompok Actinopterygii berawal dari periode Silurian sampai
awal Devonian (400 miliar tahun lalu). Kelompok ini merupakan kelompok
monofiletik dengan anggota lebih kurang sebanyak 23.000 spesies.
http://science-query.com/247740/klasifikasi-ikanpisces/
susanti, Baiq Hana, dkk. 2010. Pengantar Zoologi Vertebrata. Jakarta: Lembaga Penelitian UIN
Syarif Hidayatullah Jakarta

KELOMPOK 3
ANATOMI DAN MORFOLOGI PISCES
Awalia Maulfi
Endah Maulida
Trisniasih
Fitri Nurjannah
Adi Phati M
M. Nuruzzaman SD
Bagian tubuh ikan mulai dari anterior sampai posterior berturut-turut adalah :
1. Kepala (camput), adalah bagian tubuh mulai dari ujung mulut sampai bagian belakang
operculum
2. Tubuh (truncus), adalah bagian tubuh dari batas akhir operculum sampai anus.
3. Ekor (cauda), adalah bagian dari anus sampai bagian ujung sirip ekor terbelakang.

Bentuk luar dari organisme ini merupakan salah satu ciri yang mudah dilihat dan
diingat dalam mempelajari organisme. Adapun yang dimaksud dengan bentuk luar organisme
ini adalah bentuk tubuh, termasuk di dalamnya warna tubuh yang kelihatan dari luar. Pada
dasarnya bentuk luar dari ikan dan berbagai jenis hewan air lainnya mulai dari lahir hingga ikan
tersebut tua dapat berubah-ubah, terutama pada ikan dan hewan air lainnya yang mengalami
metamorfosis dan mengalami proses adaptasi terhadap lingkungan (habitat). Namun demikian
pada sebagian besar ikan bentuk tubuhnya relatif tetap, sehingga kalaupun terjadi perubahan,
perubahan bentuk tubuhnya relatif sangat sedikit. Bentuk tubuh pada mahluk hidup, termasuk
pada hewan air juga erat kaitannya dengan anatomi, sehingga ada baiknya sebelum melihat
anatominya; terlebih dahulu kita melihat bentuk tubuh atau penampilan (morfologi) hewan air
tersebut. Morfologi adalah bentuk tubuh (termasuk warna) yang kelihatan dari luar. Bentuk
tubuh pada mahluk hidup, termasuk pada hewan air erat kaitannya dengan anatomi, sehingga
ada baiknya sebelum melihat anatominya; terlebih dahulu kita melihat bentuk tubuh atau
penampilan (morfologi) hewan air tersebut.
Pada dasarnya morfologi dari setiap jenis hewan air yang masih dekat kekerabatanya
mempunyai kemiripan-kemiripan, seperti anatomi dan morfologi udang, kepiting dan lobster
hampir mirip. Hal yang sama juga akan kita dapati pada berbagai jenis ikan serta pada berbagai
jenis hewan lainya. Pada dasarnya kita mengenal berbagai jenis hewan air, diantaranya yang
paling umum kita kenal adalah ikan, udang, moluska, amfibi, dan sebagainya. Adapun yang
dimaksud dengan ikan adalah hewan bertulang belakang (vertebrata) yang berdarah dingin,
hidup diair, bergerak dan mempertahankan keseimbangan tubuhnya dengan menggunakan
sirip; dan bernafas dengan insang, namun selain menggunakan insang ada juga ikan yang

memiliki alat pernafasan tambahan yang fungsinya sama dengan paru-paru.

Ikan
Pada ikan dan pada hewan air lainnya pada umumnya bagian tubuh dibagi menjadi tiga bagian
yakni bagian kepala, badan dan ekor (Gambar 1), namun pada setiap jenis ikan ukuran bagianbagian tubuh tersebut berbeda-beda tergantung jenis ikannya (perhatikan morfologi ikan pada
Gambar 3) . Adapun organ-organ yang terdapat pada setiap bagian tersebut adalah:
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1007abe000000001c000000002000001847030018670300010000006c00000000000000000000
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0000001000000000000000000000000000000000400005802000069010000d40000000000000
000000000000000001c830500d23a0300460000002c00000020000000454d462b014001001c00
0000100000000210c0db010000006000000060000000460000005000000044000000454d462b
224004000c000000000000001e4009000c00000000000000244001000c0000000000000030400
20010000000040000000000803f214007000c00000000000000460000000cfe000000fe0000454
d462b08400085fcfd0000f0fd0000203b01000210c0db010000000000000000000000000000000
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6e1bc7e391d701ebca8f70242d89788da12c022c63b5d3cebb2d8b5fe822e7f8cbb81c2086f5125
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1 Pipih/kompres yakni ikan yang bertubuh pipih atau dengan kata lain lebar tubuh jauh lebih
kecil dibanding tinggi tubuh dan panjang tubuh
2 Picak/depres yakni ikan yang lebar tubuhnya jauh lebih besar dari tinggi tubuhnya
3 Cerutu/fusiform yakni ikan dengan tinggi tubuh yang hampir sama dengan lebar dan panjang
tubuhnya beberapa kali ukuran tingginya
4 Ular/sidat yakni ikan yang bentuk tubuhnya menyerupaibelut atau ular
5 Tali/filiform yakni ikan yang bentuk tubuhnya menyerupai tali
6 Pita/taeniform/flattedform yakni ikan yang bentuk tubuhnya memanjang dan tipis menyerupai
pita
7 Panah/sagittiform yakni ikan yang bentuk tubuhnya menyerupai anak panah
8 Bola/globiform yakni ikan yang bentuk tubuhnyamenyerupai bola
9 Kotak/ostraciform yakni ikan yang bentuk tubuhnya menyerupai kotak
b.Non simetri bilateral
Ikan yang non simetri bilateral diantaranya adalah ikan sebelah dan ikan lidah
2. Bentuk Mulut Ikan
010009000003a61001000000bf7f000000001610000026060f002220574d46430100000000000

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Daftar Pustaka
http://o-nlinenews.blogspot.com/2010/04/morfologi-dan-anatomi-pisces.html

KELOMPOK 4/ PD.BIOLOGI 6B
1.

Mutiara Muharram

109016100038

2.

Yunia Andriyani

109016100039

3.

Lola Novitasari

109016100044

4.

Karina Koestiarti

109016100046

5.

Dwi Septiana

6.

Muhamad Pahrudin 109016100071

109016100061

FISIOLOGI IKAN (PISCES)

A. SISTEM EKSKRESI
Sistem Ekskresi ialah sistem pembuangan proses metabolisme tubuh (berupa gas, cairan,
dan padatan melalui kulit, ginjal, dan saluran pencernaan). Sistem eksresi ikan seperti juga
pada vertebrata lain, yang mempunyai banyak fungsi antara lain untuk regulasi kadar air
tubuh, menjaga keseimbangan garam dan mengeliminasi sisa nitrogen hasil dari
metabolisme protein.
Alat pengeluaran ikan terdiri dari:
1.

Insang yang mengeluarkan CO2 dan H2O

2.

Kulit,kelenjar

kulitnya

mengeluarkan

lendir

sehingga

tubuhnya

licin

untuk memudahkan gerak di dalam air.

3.

Sepasang ginjal (sebagian besar) yang mengeluarkan urine.

Ikan beradaptasi terhadap lingkungannya dengan cara khusus. Terdapat perbedaan adaptasi
antara ikan air laut dan ikan air tawar dalam proses eksresi. Keduanya memiliki cara yang
berlawanan dalam mempertahankan keseimbangan kadar garam di dalam tubuhnya.
Air garam cenderung menyebabkan tubuh terdehidrasi, sedangkan pada kadar garam
rendah dapat menyebabkan naiknya konsentrasi garam tubuh. Ginjal ikan harus berperan
besar untuk menjaga keseimbangan garam tubuh. Beberapa ikan laut memiliki kelenjar

eksresi garam pada insang, yang berperan dalam mengeliminasi kelebihan garam. Ginjal
berfungsi untuk menyaring sesuatu yang terlarut dalam air darah dan hasilnya akan
dikeluarkan lewat korpus renalis. Tubulus yang bergulung berperan penting dalam
menjaga keseimbangan air. Hasil yang hilang pada bagian tubulus nefron, termasuk air dan
yang lain, diabsorpsi lagi ke dalam aliran darah. Korpus renalis lebih besar pada ikan air
tawar daripada ikan air laut, sehingga cairan tubuh tidak banyak keluar karena penting
untuk menjaga over dilusi (agar cairan tubuh tidak terlalu encer). Elasmobranchii, tidak
seperti kebanyakan ikan air laut, memiliki korpus renalis yang besar dan mengeluarkan air
relatif banyak, seperti pada ikan air tawar. Bangunan seperti kantung kemih pada beberapa
jenis ikan hanya untuk penampung urine sementara, dan umumnya hanya berupa perluasan
dari bagian akhir duktus ekskretori.

OSMOREGULASI
Sistem Osmoregulasi ialah sistem pengaturan keseimbangan tekanan osmotik cairan
tubuh (air dan darah) dengan tekanan osmotik habitat (perairan). Pengaturan
tekanan osmotik cairan tubuh yang layak bagi kehidupan ikan,
sehingga proses- proses fisiologis tubuhnya berfungsi normal.
Tekanan osmotik () adalah tekanan yang diberikan pada larutan
yang dapat menghentikan perpindahan molekul-molekul pelarut ke dalam
larutan melalui membran semi permeabel (proses osmosis).
Osmoregulasi dilakukan dengan berbagai cara melalui:

ginjal

kulit

membran mulut

Osmoregulasi pada ikan air tawar:

Ikan air tawar cenderung untuk menyerap air dari lingkungannya dengan cara osmosis.
Insang ikan air tawar secara aktif memasukkan garam dari lingkungan ke dalam tubuh.
Ginjal akan memompa keluar kelebihan air sebagai air seni. Ginjal mempunyai
glomeruli dalam jumlah banyak dengan diameter besar. Ini dimaksudkan untuk lebih
dapat menahan garam-garam tubuh agar tidak keluar dan sekaligus memompa air seni
sebanyak-banyaknya. Ketika cairan dari badan malpighi memasuki tubuli ginjal, glukosa
akan diserap kembali pada tubuli proximallis dan garam-garam diserap kembali pada
tubuli distal. Dinding tubuli ginjal bersifat impermiable (kedap air, tidak dapat ditembus)

terhadap air.
Osmoregulasi pada ikan air Laut:
Urine yang dihasilkan mengandung konsentrasi air yang tinggi. Ikan air laut memiliki
konsentrasi garam yang tinggi di dalam darahnya. Ikan air laut cenderung untuk
kehilangan air di dalam sel-sel tubuhnya karena proses osmosis. Untuk itu, insang ikan
air laut aktif mengeluarkan garam dari tubuhnya. Untuk mengatasi kehilangan air, ikan
minumair laut sebanyak-banyaknya. Dengan demikian berarti pula kandungan garam
akan meningkat dalam cairan tubuh. Padahal dehidrasi dicegah dengan proses ini dan
kelebihan garam harus dihilangkan. Karena ikan laut dipaksa oleh kondisi osmotik untuk
mempertahankan air, volume air seni lebih sedikit dibandingkan dengan ikan air tawar.
Tubuli ginjal mampu berfungsi sebagai penahan air. Jumlah glomeruli ikan laut
cenderung lebih sedikit dan bentuknya lebih kecil daripada ikan air tawar.
Fungsi Ginjal :

menyaring sisa-sisa proses metabolisme untuk dibuang, zat-zat yang diperlukan

tubuh diedarkan lagi melalui darah.

mengatur kekentalan urin yang dibuang untuk menjaga keseimbangan tekanan

osmotik cairan tubuh

Gambar srtuktur organ dalam ikan :

1. Hati
2. Perut
3. Usus
4. Jantung

5. Berenang kandung kemih

6. Ginjal
7. Buah pelir
8. Saluran kencing
9. Duktus eferen
10. Kandung kemih
11. Insang
Tekanan osmotik cairan tubuh berbeda antara ikan-ikan bertulang benar (Teleostei)
yang hidup di laut dengan yang hidup di perairan tawar, demikian juga dengan ikanikan bertulang rawan (Elasmobranchii), sehingga struktur dan jumlah ginjalnya juga
berbeda, demikian juga dengan sistem osmoregulasinya.
Osmoregulasi sangat penting pada hewan air karena tubuh ikan bersifat permeabel
terhadap lingkungan maupun lautan garam. Sifat fisik lingkungan yang berbeda
menyebabkan ada perbedaan proses osmoregulasi antara ikan air tawar dengan ikan air
laut. Urea merupakan produk metabolisme nitrogen, yang dikeluarkan dari tubuh ikan
berupa urin tetapi jumlahnya sedikit. Pada ikan air tawar penyerapan urea berfungsi
untuk mempertahankan tekanan osmotik dalam tubuh.
B. SISTEM REPRODUKSI
Dalam dunia perikanan, proses pembuahan ikan dikenal ada 2 macam, yakni
pembuahan internal dan eksternal. Namun demikian kebanyakan jenis ikan melakukan
pembuahan di luar.
Ikan yang melakukan pembuahan di luar disebut ikan jenis ovipar. Proses pembuahan
sel telur dan sperma berlangsung di luar tubuh ikan, dimana ikan betina tidak
mengeluarkan telur yang bercangkang namun mengeluarkan ovum yang tidak akan
berkembang lebih lanjut apabila tidak dibuahi oleh sperma. Ovum tersebut dikeluarkan
melalui ovarium melalui oviduct dan dikeluarkan melalui kloaka.
Saat akan bertelur, ikan betina mencari tempat yang rimbun oleh tumbuhan air atau di
antara bebatuan di antara air. Bersamaan dengan itu, ikan jantan juga mengeluarkan
sperma dari testis yang disalurakan melalui saluran urogenital (saluran kemih sekaligus

saluran sperma) dan keluar melalui kloaka, sehingga terjadi fertilisasi di dalam air
(fertilisasi eksternal).
C. SISTEM PENCERNAAN
Saluran pencernaan pada ikan dimulai dari rongga mulut (cavum oris). Di dalam rongga
mulut terdapat gigi-gigi kecil yang berbentuk kerucut pada geraham bawah dan lidah
pada dasar mulut yang tidak dapat digerakan serta banyak menghasilkan lendir, tetapi
tidak menghasilkan ludah (enzim). Dari rongga mulut makanan masuk ke esophagus
melalui faring yang terdapat di daerah sekitar insang.

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Esofagus berbentuk kerucut, pendek, terdapat di belakang insang, dan bila tidak dilalui
makanan lumennya menyempit. Dari kerongkongan makanan di dorong masuk ke
lambung, lambung pada umum-nya membesar, tidak jelas batasnya dengan usus. Pada

beberapa jenis ikan, terdapat tonjolan buntu untuk memperluas bidang penyerapan
makanan. Dari lambung, makanan masuk ke usus yang berupa pipa panjang berkelokkelok dan sama besarnya. Usus bermuara pada anus.
D. SISTEM TRANSPORTASI
Jantung ikan terdiri 2 ruang, yaitu 1 atrium (serambi) dan 1 ventrikel (bilik). Arah
sirkulasinya :
1. Darah dari ventrikel ke insang.
2. Pada dinding kapiler insang (pelepasan CO2 dan pengambilan O2) Aorta dorsal,
kemudian ke kapiler seluruh tubuh untuk memberikan O2 dan sari makanan serta
mengikat CO2
3. Selanjutnya darah kembali keantrium jantung melalui vena.

Seekor ikan, darah harus mengalir melalui dua hamparan kapiler selama masing-masing
sirkuit (perputaran), satu dalam insang dan kedua dikapiler sistemik. Ketika darah
mengalir melalui hamparan kepiler, tekanan darah, tekanan hidrostatik yang mendorong
darah mengalir melalui pembuluh, menurun tajam. Dengan demikian, darah yang kaya
akan oksigen dari insang mengalir ke organ-organ lain dengan sangt lambat pada ikan,
tetapi proses tersebut di bantu oleh pergerakan tubuh selama berenang.
E. SISTEM PERNAFASAN
Ikan bernapas menggunakan insang. Insang berbentuk lembaran-lembaran tipis

berwarna merah muda dan selalu lembap. Bagian terluar dari insang berhubungan
dengan air, sedang bagian dalam berhubungan erat dengan kapilerkapiler darah. Tiap
lembaran insang terdiri dari sepasang filamen dan tiap filamen mengandung banyak
lapisan tipis (lamela). Pada filamen terdapat pembuluh darah yang memiliki banyak
kapiler, sehingga memungkinkan O2 berdifusi masuk dan CO2 berdifusi keluar.
Pada ikan bertulang sejati (Osteichthyes) insangnya dilengkapi dengan tutup insang
(operkulum), sedangkan pada ikan bertulang rawan (Chondrichthyes) insangnya tidak
mempunyai tutup insang. Selain bernapas dengan insang, ada pula kelompok ikan yang
bernapas dengan gelembung udara (pulmosis), yaitu ikan paru-paru (Dipnoi). Insang
tidak hanya berfungsi sebagai alat pernapasan, tetapi juga berfungsi sebagai alat
ekskresi garam-garam, penyaring makanan, alat pertukaran ion, dan osmoregulator.
1) Sistem Pernapasan pada ikan bertulang sejati
Salah satu contoh ikan bertulang sejati yaitu ikan mas. Insang ikan mas tersimpan
dalam rongga insang yang terlindung oleh tutup insang (operkulum). Perhatikan
Gambar 7.16. Insang ikan mas terdiri dari lengkung insang yang tersusun atas tulang
rawan berwarna putih, rigi-rigi insang yang berfungsi untuk menyaring air
pernapasan yang melalui insang, dan filamen atau lembaran insang. Filamen insang
tersusun atas jaringan lunak, berbentuk sisir dan berwarna merah muda karena
mempunyai banyak pembuluh kapiler darah dan merupakan cabang dari arteri
insang. Di tempat inilah pertukaran gas CO2 dan O2 berlangsung.
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fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7ff
f7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7
fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7ff
f7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7
fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7ff
f7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fde7bff7fff
7fff7fde7f4a29ce39b55aff7f7c73d65ac6181963ff7f4b29f041ff7f5b6bc618f85eff7fbd73e81c94
52bd77396708258b31ff7f5a6be814f95aff7fef414931dc7fdd7f0729724
Gas O2 diambil dari gas O2 yang larut dalam air melalui insang secara difusi. Dari

insang, O2 diangkut darah melalui pembuluh darah ke seluruh jaringan tubuh. Dari
jaringan tubuh, gas CO2 diangkut darah menuju jantung. Dari jantung menuju insang
untuk melakukan pertukaran gas. Proses ini terjadi secara terus-menerus dan
berulang-ulang.
Mekanisme pernapasan ikan bertulang sejati dilakukan melalui mekanisme inspirasi
dan ekspirasi. Perhatikan Gambar 7.17.
0100090000038e0c0500000007de020000001610000026060f002220574d46430100000000000
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0000001000000000000000000000000000000000400005802000069010000d40000000000000
000000000000000001c830500d23a0300460000002c00000020000000454d462b014001001c00
0000100000000210c0db01000000600000006000000046000000887f00007c7f0000454d462b2
24004000c000000000000001e4009000c00000000000000244001000c00000000000000304002
0010000000040000000000803f214007000c0000000000000008400005d47e0000c87e0000021
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232323232323232323232323232323232323232323232323232323232ffc0001108020700f103
012200021101031101ffc4001f000001050101010101010000000000000000010203040506070
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a) Fase inspirasi ikan

Gerakan tutup insang ke samping dan selaput tutup insang tetap menempel pada
tubuh mengakibatkan rongga mulut bertambah besar, sebaliknya celah belakang
insang tertutup. Akibatnya, tekanan udara dalam rongga mulut lebih kecil daripada
tekanan udara luar. Celah mulut membuka sehingga terjadi aliran air ke dalam
rongga mulut.
b) Fase ekspirasi ikan
Setelah air masuk ke dalam rongga mulut, celah mulut menutup. Insang kembali ke
kedudukan semula diikuti membukanya celah insang. Air dalam mulut mengalir
melalui celah-celah insang dan menyentuh lembaran-lembaran insang. Pada tempat
ini terjadi pertukaran udara pernapasan. Darah melepaskan CO2 ke dalam air dan
mengikat O2 dari air.
Insang tidak saja berfungsi sebagai alat pernapasan tetapi dapat pula berfungsi
sebagai alat ekskresi garam-garam, penyaring makanan, alat pertukaran ion,
dan osmoregulator. Beberapa jenis ikan mempunyai labirin yang merupakan
perluasan ke atas dari insang dan membentuk lipatan-lipatan sehingga merupakan
rongga-rongga tidak teratur. Labirin ini berfungsi menyimpan cadangan O2 sehingga

ikan tahan pada kondisi yang kekurangan O2. Contoh ikan yang mempunyai labirin
adalah: ikan gabus dan ikan lele. Untuk menyimpan cadangan O2, selain dengan
labirin, ikan mempunyai gelembung renang yang terletak di dekat punggung.
KELOMPOK 4/ PD.BIOLOGI 6B
7.

Mutiara Muharram

109016100038

8.

Yunia Andriyani

109016100039

9.

Lola Novitasari

109016100044

10. Karina Koestiarti

11. Dwi Septiana

109016100046
109016100061

12. Muhamad Pahrudin 109016100071

FISIOLOGI IKAN (PISCES)

F. SISTEM EKSKRESI
Sistem Ekskresi ialah sistem pembuangan proses metabolisme tubuh (berupa gas, cairan,
dan padatan melalui kulit, ginjal, dan saluran pencernaan). Sistem eksresi ikan seperti juga
pada vertebrata lain, yang mempunyai banyak fungsi antara lain untuk regulasi kadar air
tubuh, menjaga keseimbangan garam dan mengeliminasi sisa nitrogen hasil dari
metabolisme protein.
Alat pengeluaran ikan terdiri dari:
4.

Insang yang mengeluarkan CO2 dan H2O

5.

Kulit,kelenjar

kulitnya

mengeluarkan

lendir

sehingga

tubuhnya

licin

untuk memudahkan gerak di dalam air.

6.

Sepasang ginjal (sebagian besar) yang mengeluarkan urine.

Ikan beradaptasi terhadap lingkungannya dengan cara khusus. Terdapat perbedaan adaptasi
antara ikan air laut dan ikan air tawar dalam proses eksresi. Keduanya memiliki cara yang
berlawanan dalam mempertahankan keseimbangan kadar garam di dalam tubuhnya.
Air garam cenderung menyebabkan tubuh terdehidrasi, sedangkan pada kadar garam
rendah dapat menyebabkan naiknya konsentrasi garam tubuh. Ginjal ikan harus berperan
besar untuk menjaga keseimbangan garam tubuh. Beberapa ikan laut memiliki kelenjar

eksresi garam pada insang, yang berperan dalam mengeliminasi kelebihan garam. Ginjal
berfungsi untuk menyaring sesuatu yang terlarut dalam air darah dan hasilnya akan
dikeluarkan lewat korpus renalis. Tubulus yang bergulung berperan penting dalam
menjaga keseimbangan air. Hasil yang hilang pada bagian tubulus nefron, termasuk air dan
yang lain, diabsorpsi lagi ke dalam aliran darah. Korpus renalis lebih besar pada ikan air
tawar daripada ikan air laut, sehingga cairan tubuh tidak banyak keluar karena penting
untuk menjaga over dilusi (agar cairan tubuh tidak terlalu encer). Elasmobranchii, tidak
seperti kebanyakan ikan air laut, memiliki korpus renalis yang besar dan mengeluarkan air
relatif banyak, seperti pada ikan air tawar. Bangunan seperti kantung kemih pada beberapa
jenis ikan hanya untuk penampung urine sementara, dan umumnya hanya berupa perluasan
dari bagian akhir duktus ekskretori.

OSMOREGULASI
Sistem Osmoregulasi ialah sistem pengaturan keseimbangan tekanan osmotik cairan
tubuh (air dan darah) dengan tekanan osmotik habitat (perairan). Pengaturan tekanan
osmotik cairan tubuh yang layak bagi kehidupan ikan, sehingga proses- proses
fisiologis tubuhnya berfungsi normal. Tekanan osmotik () adalah
tekanan yang diberikan pada larutan yang dapat menghentikan
perpindahan molekul-molekul pelarut ke dalam larutan melalui
membran semi permeabel (proses osmosis).
Osmoregulasi dilakukan dengan berbagai cara melalui:

ginjal

kulit

membran mulut

Osmoregulasi pada ikan air tawar:

Ikan air tawar cenderung untuk menyerap air dari lingkungannya dengan cara osmosis.
Insang ikan air tawar secara aktif memasukkan garam dari lingkungan ke dalam tubuh.
Ginjal akan memompa keluar kelebihan air sebagai air seni. Ginjal mempunyai
glomeruli dalam jumlah banyak dengan diameter besar. Ini dimaksudkan untuk lebih
dapat menahan garam-garam tubuh agar tidak keluar dan sekaligus memompa air seni
sebanyak-banyaknya. Ketika cairan dari badan malpighi memasuki tubuli ginjal, glukosa
akan diserap kembali pada tubuli proximallis dan garam-garam diserap kembali pada
tubuli distal. Dinding tubuli ginjal bersifat impermiable (kedap air, tidak dapat ditembus)

terhadap air.
Osmoregulasi pada ikan air Laut:
Urine yang dihasilkan mengandung konsentrasi air yang tinggi. Ikan air laut memiliki
konsentrasi garam yang tinggi di dalam darahnya. Ikan air laut cenderung untuk
kehilangan air di dalam sel-sel tubuhnya karena proses osmosis. Untuk itu, insang ikan
air laut aktif mengeluarkan garam dari tubuhnya. Untuk mengatasi kehilangan air, ikan
minumair laut sebanyak-banyaknya. Dengan demikian berarti pula kandungan garam
akan meningkat dalam cairan tubuh. Padahal dehidrasi dicegah dengan proses ini dan
kelebihan garam harus dihilangkan. Karena ikan laut dipaksa oleh kondisi osmotik untuk
mempertahankan air, volume air seni lebih sedikit dibandingkan dengan ikan air tawar.
Tubuli ginjal mampu berfungsi sebagai penahan air. Jumlah glomeruli ikan laut
cenderung lebih sedikit dan bentuknya lebih kecil daripada ikan air tawar.
Fungsi Ginjal :

menyaring sisa-sisa proses metabolisme untuk dibuang, zat-zat yang diperlukan

tubuh diedarkan lagi melalui darah.

mengatur kekentalan urin yang dibuang untuk menjaga keseimbangan tekanan

osmotik cairan tubuh

Gambar srtuktur organ dalam ikan :

12. Hati
13. Perut
14. Usus
15. Jantung

16. Berenang kandung kemih

17. Ginjal
18. Buah pelir
19. Saluran kencing
20. Duktus eferen
21. Kandung kemih
22. Insang
Tekanan osmotik cairan tubuh berbeda antara ikan-ikan bertulang benar (Teleostei)
yang hidup di laut dengan yang hidup di perairan tawar, demikian juga dengan ikanikan bertulang rawan (Elasmobranchii), sehingga struktur dan jumlah ginjalnya juga
berbeda, demikian juga dengan sistem osmoregulasinya.
Osmoregulasi sangat penting pada hewan air karena tubuh ikan bersifat permeabel
terhadap lingkungan maupun lautan garam. Sifat fisik lingkungan yang berbeda
menyebabkan ada perbedaan proses osmoregulasi antara ikan air tawar dengan ikan air
laut. Urea merupakan produk metabolisme nitrogen, yang dikeluarkan dari tubuh ikan
berupa urin tetapi jumlahnya sedikit. Pada ikan air tawar penyerapan urea berfungsi
untuk mempertahankan tekanan osmotik dalam tubuh.
G. SISTEM REPRODUKSI
Dalam dunia perikanan, proses pembuahan ikan dikenal ada 2 macam, yakni
pembuahan internal dan eksternal. Namun demikian kebanyakan jenis ikan melakukan
pembuahan di luar.
Ikan yang melakukan pembuahan di luar disebut ikan jenis ovipar. Proses pembuahan
sel telur dan sperma berlangsung di luar tubuh ikan, dimana ikan betina tidak
mengeluarkan telur yang bercangkang namun mengeluarkan ovum yang tidak akan
berkembang lebih lanjut apabila tidak dibuahi oleh sperma. Ovum tersebut dikeluarkan
melalui ovarium melalui oviduct dan dikeluarkan melalui kloaka.
Saat akan bertelur, ikan betina mencari tempat yang rimbun oleh tumbuhan air atau di
antara bebatuan di antara air. Bersamaan dengan itu, ikan jantan juga mengeluarkan
sperma dari testis yang disalurakan melalui saluran urogenital (saluran kemih sekaligus

saluran sperma) dan keluar melalui kloaka, sehingga terjadi fertilisasi di dalam air
(fertilisasi eksternal).
H. SISTEM PENCERNAAN
Saluran pencernaan pada ikan dimulai dari rongga mulut (cavum oris). Di dalam rongga
mulut terdapat gigi-gigi kecil yang berbentuk kerucut pada geraham bawah dan lidah
pada dasar mulut yang tidak dapat digerakan serta banyak menghasilkan lendir, tetapi
tidak menghasilkan ludah (enzim). Dari rongga mulut makanan masuk ke esophagus
melalui faring yang terdapat di daerah sekitar insang.

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fff7f9c738c311042ad3529250821f75e3967a51

Esofagus berbentuk kerucut, pendek, terdapat di belakang insang, dan bila tidak dilalui
makanan lumennya menyempit. Dari kerongkongan makanan di dorong masuk ke
lambung, lambung pada umum-nya membesar, tidak jelas batasnya dengan usus. Pada

beberapa jenis ikan, terdapat tonjolan buntu untuk memperluas bidang penyerapan
makanan. Dari lambung, makanan masuk ke usus yang berupa pipa panjang berkelokkelok dan sama besarnya. Usus bermuara pada anus.
I. SISTEM TRANSPORTASI
Jantung ikan terdiri 2 ruang, yaitu 1 atrium (serambi) dan 1 ventrikel (bilik). Arah
sirkulasinya :
4. Darah dari ventrikel ke insang.
5. Pada dinding kapiler insang (pelepasan CO2 dan pengambilan O2) Aorta dorsal,
kemudian ke kapiler seluruh tubuh untuk memberikan O2 dan sari makanan serta
mengikat CO2
6. Selanjutnya darah kembali keantrium jantung melalui vena.

Seekor ikan, darah harus mengalir melalui dua hamparan kapiler selama masing-masing
sirkuit (perputaran), satu dalam insang dan kedua dikapiler sistemik. Ketika darah
mengalir melalui hamparan kepiler, tekanan darah, tekanan hidrostatik yang mendorong
darah mengalir melalui pembuluh, menurun tajam. Dengan demikian, darah yang kaya
akan oksigen dari insang mengalir ke organ-organ lain dengan sangt lambat pada ikan,
tetapi proses tersebut di bantu oleh pergerakan tubuh selama berenang.
J. SISTEM PERNAFASAN
Ikan bernapas menggunakan insang. Insang berbentuk lembaran-lembaran tipis

berwarna merah muda dan selalu lembap. Bagian terluar dari insang berhubungan
dengan air, sedang bagian dalam berhubungan erat dengan kapilerkapiler darah. Tiap
lembaran insang terdiri dari sepasang filamen dan tiap filamen mengandung banyak
lapisan tipis (lamela). Pada filamen terdapat pembuluh darah yang memiliki banyak
kapiler, sehingga memungkinkan O2 berdifusi masuk dan CO2 berdifusi keluar.
Pada ikan bertulang sejati (Osteichthyes) insangnya dilengkapi dengan tutup insang
(operkulum), sedangkan pada ikan bertulang rawan (Chondrichthyes) insangnya tidak
mempunyai tutup insang. Selain bernapas dengan insang, ada pula kelompok ikan yang
bernapas dengan gelembung udara (pulmosis), yaitu ikan paru-paru (Dipnoi). Insang
tidak hanya berfungsi sebagai alat pernapasan, tetapi juga berfungsi sebagai alat
ekskresi garam-garam, penyaring makanan, alat pertukaran ion, dan osmoregulator.
1) Sistem Pernapasan pada ikan bertulang sejati
Salah satu contoh ikan bertulang sejati yaitu ikan mas. Insang ikan mas tersimpan
dalam rongga insang yang terlindung oleh tutup insang (operkulum). Perhatikan
Gambar 7.16. Insang ikan mas terdiri dari lengkung insang yang tersusun atas tulang
rawan berwarna putih, rigi-rigi insang yang berfungsi untuk menyaring air
pernapasan yang melalui insang, dan filamen atau lembaran insang. Filamen insang
tersusun atas jaringan lunak, berbentuk sisir dan berwarna merah muda karena
mempunyai banyak pembuluh kapiler darah dan merupakan cabang dari arteri
insang. Di tempat inilah pertukaran gas CO2 dan O2 berlangsung.
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fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7fff7ff
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52bd77396708258b31ff7f5a6be814f95aff7fef414931dc7fdd7f0729724
Gas O2 diambil dari gas O2 yang larut dalam air melalui insang secara difusi. Dari

insang, O2 diangkut darah melalui pembuluh darah ke seluruh jaringan tubuh. Dari
jaringan tubuh, gas CO2 diangkut darah menuju jantung. Dari jantung menuju insang
untuk melakukan pertukaran gas. Proses ini terjadi secara terus-menerus dan
berulang-ulang.
Mekanisme pernapasan ikan bertulang sejati dilakukan melalui mekanisme inspirasi
dan ekspirasi. Perhatikan Gambar 7.17.
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a) Fase inspirasi ikan

Gerakan tutup insang ke samping dan selaput tutup insang tetap menempel pada
tubuh mengakibatkan rongga mulut bertambah besar, sebaliknya celah belakang
insang tertutup. Akibatnya, tekanan udara dalam rongga mulut lebih kecil daripada
tekanan udara luar. Celah mulut membuka sehingga terjadi aliran air ke dalam
rongga mulut.
b) Fase ekspirasi ikan
Setelah air masuk ke dalam rongga mulut, celah mulut menutup. Insang kembali ke
kedudukan semula diikuti membukanya celah insang. Air dalam mulut mengalir
melalui celah-celah insang dan menyentuh lembaran-lembaran insang. Pada tempat
ini terjadi pertukaran udara pernapasan. Darah melepaskan CO2 ke dalam air dan
mengikat O2 dari air.
Insang tidak saja berfungsi sebagai alat pernapasan tetapi dapat pula berfungsi
sebagai alat ekskresi garam-garam, penyaring makanan, alat pertukaran ion,
dan osmoregulator. Beberapa jenis ikan mempunyai labirin yang merupakan
perluasan ke atas dari insang dan membentuk lipatan-lipatan sehingga merupakan
rongga-rongga tidak teratur. Labirin ini berfungsi menyimpan cadangan O2 sehingga

ikan tahan pada kondisi yang kekurangan O2. Contoh ikan yang mempunyai labirin
adalah: ikan gabus dan ikan lele. Untuk menyimpan cadangan O2, selain dengan
labirin, ikan mempunyai gelembung renang yang terletak di dekat punggung.